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Rattan's beardtongue, Rattan's penstemon

Habit Herbs.
Stems

ascending to erect, (15–)30–120 cm, glabrous, sometimes glabrous proximally, glandular-pubescent distally, not glaucous.

Leaves

basal and cauline, not leathery, glabrous;

basal and proximal cauline 30–250 × 7–50 mm, blade elliptic to lanceolate, base tapered, margins serrate to dentate, apex obtuse to acute;

cauline 2–5 pairs, short-petiolate or sessile, (10–)20–75(–110) × 5–30(–38) mm, blade oblong to triangular-ovate, base truncate to cordate-clasping distally, margins serrate to dentate, apex obtuse to acute.

Thyrses

interrupted, cylindric, 2–24 cm, axis glandular-pubescent, verticillasters 2–7, cymes 3–11-flowered, 2 per node;

proximal bracts triangular to lanceolate, 6–50 × 1–22 mm, margins entire or ± serrate;

peduncles and pedicels ascending to erect, glandular-pubescent.

Flowers

calyx lobes ovate to ovate-oblong or lanceolate, 4–10 × (1–)1.5–2.5 mm, glandular-pubescent;

corolla lavender to purple or violet, with reddish purple nectar guides, ventricose to ventricose-ampliate, 20–30 mm, glandular-pubescent externally, sparsely to densely white-lanate internally, tube 6–7 mm, throat abruptly inflated, 8–10 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, subexplanate, 1.2–1.5 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate;

staminode 21–26 mm, exserted, 0.3–0.4 mm diam., tip straight to recurved, distal 6–10 mm sparsely to moderately lanate, hairs white or yellow, to 3 mm;

style 16–19 mm.

Capsules

5–9 × 3.5–5 mm, glabrous.

2n

= 16.

Penstemon rattanii

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Penstemon rattanii comprises two morphologically distinct and geographically allopatric varieties confined to the Coast Range of California and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes usually longer than capsules, lanceolate, sometimes ovate, (5.5–)6.5–10 mm, apices acute to acuminate, sometimes obtuse; nw California, sw Oregon.
P. rattanii var. rattanii
1. Calyx lobes usually shorter than capsules, ovate-oblong, 4–7 mm, apices obtuse to acute; wc California.
P. rattanii var. kleei
Source FNA vol. 17, p. 219.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. rattanii var. kleei, P. rattanii var. rattanii
Name authority A. Gray: Proc. Amer. Acad. Arts 15: 50. (1879)
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