Rattan's beardtongue, Rattan's penstemon
|
Degener's beardtongue
|
Herbs. |
Herbs. |
ascending to erect, (15–)30–120 cm, glabrous, sometimes glabrous proximally, glandular-pubescent distally, not glaucous. |
ascending to erect, 25–40 cm, retrorsely hairy proximally, glandular-pubescent distally, not glaucous. |
basal and cauline, not leathery, glabrous; basal and proximal cauline 30–250 × 7–50 mm, blade elliptic to lanceolate, base tapered, margins serrate to dentate, apex obtuse to acute; cauline 2–5 pairs, short-petiolate or sessile, (10–)20–75(–110) × 5–30(–38) mm, blade oblong to triangular-ovate, base truncate to cordate-clasping distally, margins serrate to dentate, apex obtuse to acute. |
basal and cauline, basal sometimes withering by anthesis, not leathery, glabrous or retrorsely hairy proximally; basal and proximal cauline 25–70(–85) × 5–23 mm, blade ovate to lanceolate, base tapered, margins entire, apex acute; cauline 3–6 pairs, petiolate or sessile, 20–90(–110) × 2–18 mm, blade oblanceolate to lanceolate, base tapered, margins entire, apex acute. |
interrupted, cylindric, 2–24 cm, axis glandular-pubescent, verticillasters 2–7, cymes 3–11-flowered, 2 per node; proximal bracts triangular to lanceolate, 6–50 × 1–22 mm, margins entire or ± serrate; peduncles and pedicels ascending to erect, glandular-pubescent. |
interrupted, narrowly conic, 3–10 cm, axis sparsely glandular-pubescent, verticillasters 3–6, cymes 2–10-flowered, 2 per node; proximal bracts lanceolate to linear, 3–40 × 0.5–4 mm, margins entire; peduncles and pedicels ascending to erect, sparsely glandular-pubescent. |
calyx lobes ovate to ovate-oblong or lanceolate, 4–10 × (1–)1.5–2.5 mm, glandular-pubescent; corolla lavender to purple or violet, with reddish purple nectar guides, ventricose to ventricose-ampliate, 20–30 mm, glandular-pubescent externally, sparsely to densely white-lanate internally, tube 6–7 mm, throat abruptly inflated, 8–10 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate, 1.2–1.5 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode 21–26 mm, exserted, 0.3–0.4 mm diam., tip straight to recurved, distal 6–10 mm sparsely to moderately lanate, hairs white or yellow, to 3 mm; style 16–19 mm. |
calyx lobes ovate to lanceolate, 3.5–7 × 1–2 mm, glandular-pubescent; corolla blue to light violet, with reddish purple nectar guides, funnelform, 14–19 mm, glandular-pubescent or glabrescent externally, glabrous or sparsely white- or yellow-villous internally abaxially, tube 5–6 mm, throat gradually inflated, 4–9 mm diam., slightly 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, 0.9–1.1 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 15–16 mm, included, 0.3–0.4 mm diam., tip straight to slightly recurved, distal 6–10 mm pilose, hairs golden, to 1.5 mm; style 13–16 mm. |
5–9 × 3.5–5 mm, glabrous. |
7–10 × 2.5–4 mm, glabrous. |
= 16. |
|
|
|
|
|
|
Flowering Jun–Aug. |
|
Pine-juniper woodlands, ponderosa pine parklands, montane grasslands. |
|
1800–2900 m. (5900–9500 ft.) |
CA; OR
|
CO |
Varieties 2 (2 in the flora). Penstemon rattanii comprises two morphologically distinct and geographically allopatric varieties confined to the Coast Range of California and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon degeneri is known from the Arkansas River Canyon from near Salida to Cañon City, and to near Wet Mountain Valley. Most populations are in Fremont County (B. L. Beatty et al. 2004); the species is also known from Chaffee and Custer counties. Typical, large-leaved forms are unlikely to be confused with any other species in the area, but narrow-leaved forms can be mistaken for P. griffinii. F. S. Crosswhite (1965b) described the corollas of P. degeneri as bearded at the orifices with light yellow hairs. In most populations, the abaxial limbs and throats are sparsely yellow-lanate; however, pubescence can vary greatly within populations, from sparsely white-lanate only on the limbs to moderately yellow-lanate on the limbs and throats. When the throats are bearded in P. degeneri, the hairs extend 1–3 mm into the throats; in P. griffinii they extend deeper into the throats, usually 5–8 mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Calyx lobes usually longer than capsules, lanceolate, sometimes ovate, (5.5–)6.5–10 mm, apices acute to acuminate, sometimes obtuse; nw California, sw Oregon. | var. rattanii |
1. Calyx lobes usually shorter than capsules, ovate-oblong, 4–7 mm, apices obtuse to acute; wc California. | var. kleei |
|
|
FNA vol. 17, p. 219. |
FNA vol. 17, p. 199. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
|
|
A. Gray: Proc. Amer. Acad. Arts 15: 50. (1879) — (as Pentstemon rattani) |
Crosswhite: Amer. Midl. Naturalist 74: 434, fig. 4. (1965) |
| |