Rattan's beardtongue, Rattan's penstemon
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Gibbens' beardtongue
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Herbs. |
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ascending to erect, (15–)30–120 cm, glabrous, sometimes glabrous proximally, glandular-pubescent distally, not glaucous. |
ascending to erect, 10–37 cm, pubescent to retrorsely hairy, sometimes glabrous proximally, glandular-pubescent distally, not glaucous. |
basal and cauline, not leathery, glabrous; basal and proximal cauline 30–250 × 7–50 mm, blade elliptic to lanceolate, base tapered, margins serrate to dentate, apex obtuse to acute; cauline 2–5 pairs, short-petiolate or sessile, (10–)20–75(–110) × 5–30(–38) mm, blade oblong to triangular-ovate, base truncate to cordate-clasping distally, margins serrate to dentate, apex obtuse to acute. |
basal and cauline, or basal absent or reduced, not leathery, proximals glabrous or puberulent to scabrous, distals puberulent or scabrous to glandular-pubescent, not glaucous; basal and proximal cauline 15–90 × 2–7(–8) mm, blade oblanceolate to linear, base tapered, margins entire, apex obtuse to acute; cauline 6–10 pairs, short-petiolate or sessile, 9–68 × 1–5 mm, blade oblanceolate to linear, base tapered to truncate, apex obtuse to acute. |
interrupted, cylindric, 2–24 cm, axis glandular-pubescent, verticillasters 2–7, cymes 3–11-flowered, 2 per node; proximal bracts triangular to lanceolate, 6–50 × 1–22 mm, margins entire or ± serrate; peduncles and pedicels ascending to erect, glandular-pubescent. |
interrupted, secund, (2–)5–14 cm, axis glandular-pubescent, verticillasters (2–)5–8, cymes 1–3-flowered, 1 or 2 per node; proximal bracts lanceolate, proximals 8–43 × 1–3 mm; peduncles and pedicels glandular-pubescent. |
calyx lobes ovate to ovate-oblong or lanceolate, 4–10 × (1–)1.5–2.5 mm, glandular-pubescent; corolla lavender to purple or violet, with reddish purple nectar guides, ventricose to ventricose-ampliate, 20–30 mm, glandular-pubescent externally, sparsely to densely white-lanate internally, tube 6–7 mm, throat abruptly inflated, 8–10 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate, 1.2–1.5 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode 21–26 mm, exserted, 0.3–0.4 mm diam., tip straight to recurved, distal 6–10 mm sparsely to moderately lanate, hairs white or yellow, to 3 mm; style 16–19 mm. |
calyx lobes ovate to lanceolate, 3.5–7(–8) × 1.8–2.5 mm, glandular-pubescent; corolla lavender to light blue or light violet, with faint reddish purple nectar guides, funnelform, (15–)16–18(–20) mm, glandular-pubescent externally, sparsely to moderately glandular-pubescent or glandular internally abaxially, tube 5.5–6.5 mm, throat gradually inflated, not constricted at orifice, 5–6 mm diam., rounded abaxially; stamens: longer pair reaching orifice, pollen sacs divergent, navicular, 1–1.4 mm, dehiscing incompletely, proximal 1/5 indehiscent, connective not splitting, sides moderately hirsute, hairs white, to 0.7 mm, sutures denticulate, teeth to 0.1 mm; staminode 8–10 mm, included, 0.6–0.7 mm diam., tip recurved, distal 1–3 mm sparsely pilose or lanate, hairs yellow or whitish, to 1 mm; style 11–16 mm. |
5–9 × 3.5–5 mm, glabrous. |
5–8 × 3.5–5 mm. |
= 16. |
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Flowering Jun–Sep. |
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Barren hills, pinyon-juniper woodlands, sagebrush and greasewood-saltbush shrublands. |
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1700–2300 m. (5600–7500 ft.) |
CA; OR
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CO; UT; WY |
Varieties 2 (2 in the flora). Penstemon rattanii comprises two morphologically distinct and geographically allopatric varieties confined to the Coast Range of California and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon gibbensii is known from fewer than ten populations in Moffat and Rio Blanco counties, Colorado, Daggett County, Utah, and Carbon and Sweetwater counties, Wyoming (S. L. O’Kane 1988; E. C. Neese and N. D. Atwood 2003). Plants occur on shales or sandstones of the Browns Park Formation and Green River Formation (B. L. Heidel 2009). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Calyx lobes usually longer than capsules, lanceolate, sometimes ovate, (5.5–)6.5–10 mm, apices acute to acuminate, sometimes obtuse; nw California, sw Oregon. | var. rattanii |
1. Calyx lobes usually shorter than capsules, ovate-oblong, 4–7 mm, apices obtuse to acute; wc California. | var. kleei |
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FNA vol. 17, p. 219. |
FNA vol. 17, p. 167. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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A. Gray: Proc. Amer. Acad. Arts 15: 50. (1879) — (as Pentstemon rattani) |
Dorn: Brittonia 34: 334. (1982) |
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