Rattan's beardtongue, Rattan's penstemon
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Chelan beardtongue, Chelan penstemon, Chilean beardtongue
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Herbs. |
Herbs. |
ascending to erect, (15–)30–120 cm, glabrous, sometimes glabrous proximally, glandular-pubescent distally, not glaucous. |
ascending or erect, 8–40(–60) cm, glabrate, retrorsely hairy, or glandular-pubescent, not glaucous. |
basal and cauline, not leathery, glabrous; basal and proximal cauline 30–250 × 7–50 mm, blade elliptic to lanceolate, base tapered, margins serrate to dentate, apex obtuse to acute; cauline 2–5 pairs, short-petiolate or sessile, (10–)20–75(–110) × 5–30(–38) mm, blade oblong to triangular-ovate, base truncate to cordate-clasping distally, margins serrate to dentate, apex obtuse to acute. |
basal and cauline, not leathery, retrorsely hairy or pubescent, sometimes also glandular-pubescent; basal and proximal cauline 25–100(–150) × (4–)9–20(–34) mm, blade elliptic to ovate, base tapered, margins ± serrate to dentate, apex obtuse to acute; cauline 2–5 pairs, sessile, 11–60 × 5–25(–30) mm, blade triangular-ovate to lanceolate, base truncate to cordate-clasping, margins serrate to dentate, apex acute, rarely obtuse. |
interrupted, cylindric, 2–24 cm, axis glandular-pubescent, verticillasters 2–7, cymes 3–11-flowered, 2 per node; proximal bracts triangular to lanceolate, 6–50 × 1–22 mm, margins entire or ± serrate; peduncles and pedicels ascending to erect, glandular-pubescent. |
interrupted, cylindric, 4–28 cm, axis densely glandular-pubescent, rarely sparsely glandular-pubescent, verticillasters 3–8, cymes 3–11-flowered, 2 per node; proximal bracts ovate to lanceolate, 8–58 × 4–25 mm, margins serrate, sometimes entire; peduncles and pedicels ascending to erect, densely glandular-pubescent. |
calyx lobes ovate to ovate-oblong or lanceolate, 4–10 × (1–)1.5–2.5 mm, glandular-pubescent; corolla lavender to purple or violet, with reddish purple nectar guides, ventricose to ventricose-ampliate, 20–30 mm, glandular-pubescent externally, sparsely to densely white-lanate internally, tube 6–7 mm, throat abruptly inflated, 8–10 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate, 1.2–1.5 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode 21–26 mm, exserted, 0.3–0.4 mm diam., tip straight to recurved, distal 6–10 mm sparsely to moderately lanate, hairs white or yellow, to 3 mm; style 16–19 mm. |
calyx lobes ovate to lanceolate, 2.5–4.5(–6) × 0.9–1.5 mm, glandular-pubescent; corolla blue to lavender, purple, or violet, with purple nectar guides, funnelform, 11–16 mm, glandular-pubescent externally, sparsely white-villous internally abaxially, tube 4–5 mm, throat gradually inflated, 3–6 mm diam., rounded abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, navicular to subexplanate, 0.6–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 7–9 mm, included or slightly exserted, 0.2–0.3 mm diam., tip recurved, distal 0.5–1 mm densely pilose, hairs yellow, to 0.8 mm; style 8–10 mm. |
5–9 × 3.5–5 mm, glabrous. |
4–7 × 3–4 mm, glabrous. |
= 16. |
= 16. |
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Flowering Apr–Jul. |
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Open grassy, sandy, gravelly, and rocky slopes, pine and pine-fir woodlands, sagebrush shrublands. |
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300–2000 m. (1000–6600 ft.) |
CA; OR
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WA; BC
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Varieties 2 (2 in the flora). Penstemon rattanii comprises two morphologically distinct and geographically allopatric varieties confined to the Coast Range of California and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon pruinosus occurs along the eastern slope of the Cascade Range in Chelan, Douglas, Grant, and Kittitas counties, Washington, east into the scablands of Adams and Franklin counties, Washington, and north into extreme southern British Columbia. A blue dye is made from the flowers of Penstemon pruinosus by the Okanagan-Coville tribe of northeastern Washington (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Calyx lobes usually longer than capsules, lanceolate, sometimes ovate, (5.5–)6.5–10 mm, apices acute to acuminate, sometimes obtuse; nw California, sw Oregon. | var. rattanii |
1. Calyx lobes usually shorter than capsules, ovate-oblong, 4–7 mm, apices obtuse to acute; wc California. | var. kleei |
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FNA vol. 17, p. 219. |
FNA vol. 17, p. 218. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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A. Gray: Proc. Amer. Acad. Arts 15: 50. (1879) — (as Pentstemon rattani) |
Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1280. (1829) — (as Pentstemon pruinosum) |
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