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buck-horn cholla

Habit Trees or shrubs, sparsely to densely branched, 1–2(–4) m. Stem segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm; tubercles prominent, narrow to broad, 1.5–4.5 cm; areoles elliptic to subcircular, 4–5.5 × 4–5 mm; wool white or yellowish to tan, aging gray-black.
Spines

6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles;

major abaxial spines deflexed to divergent, subterete to flattened;

major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm;

sheaths white to grayish with yellow to golden tips, baggy to tight fitting.

Glochids

in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm.

Flowers

inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate;

filaments red;

anthers yellow;

style and stigma lobes white to light green.

Fruits

tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus;

basal tubercles longest;

areoles 12–30.

Seeds

pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions;

girdle smooth.

Cylindropuntia acanthocarpa

Distribution
from FNA
AZ; CA; NV; UT; n Mexico
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[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tubercles 2.5-4.5 cm, broad; stem segments gray-green; spines of adjacent areoles not or barely interlaced, 6-11(-14) per areole; fruits bearing few spines or sometimes spineless
var. thornbe
1. Tubercles (1-)1.5-2.5(-3) cm, broad to narrow; stem segments green to yellow-green; spines of adjacent areoles interlaced; fruits densely spiny, sometimes burlike
→ 2
2. Abaxial 5-7 spines deflexed, usually subparallel, whitish to light brown; tubercles broad, oval; shrubs with few, ascending branches
var. acantho
2. Abaxial (14-)18-30+ spines deflexed to divergent, spreading, pale yellow to red-brown; tubercles narrowly elliptic; shrubs with few to many, spreading branches
→ 3
3. Branches usually at acute angles; stem segments yellowish green, usually long, commonly 15-30 cm; spines 18-30+, yellow to tan, sheaths baggy; inner tepals yellow
var. colorad
3. Branches usually at obtuse angles; stem segments green to dark green, usually short, commonly 8 -20 cm; spines 14-18(-23) per areole, yellow or red-brown, sheaths tight; inner tepals yellow to bronze to brick red
var. major
Source FNA vol. 4.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Subordinate taxa
C. acanthocarpa var. acantho, C. acanthocarpa var. colorad, C. acanthocarpa var. major, C. acanthocarpa var. thornbe
Synonyms Opuntia acanthocarpa
Name authority (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935)
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