The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

buck-horn cholla

Arizona pencil cholla, pencil cholla

Habit Trees or shrubs, sparsely to densely branched, 1–2(–4) m. Stem segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm; tubercles prominent, narrow to broad, 1.5–4.5 cm; areoles elliptic to subcircular, 4–5.5 × 4–5 mm; wool white or yellowish to tan, aging gray-black. Trees or shrubs, densely branched, 0.5–3 m. Stem segments green, sometimes purple tinged, 6–10.5 × 0.5–1.3 cm; tubercles narrowly elongate, appearing as wrinkles when dry, 1–2 cm; areoles circular, 2–3 mm in diam.; wool tan to brown, aging gray to black.
Spines

6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles;

major abaxial spines deflexed to divergent, subterete to flattened;

major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm;

sheaths white to grayish with yellow to golden tips, baggy to tight fitting.

0–2(–3) per areole, sparsely distributed along stem, usually deflexed, pale yellow or red-brown aging black, stout, the longest 0.8–3.5(–5) cm;

sheaths loose fitting, yellowish brown.

Glochids

in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm.

in adaxial tuft and marginal, encircling areole, pale yellow.

Flowers

inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate;

filaments red;

anthers yellow;

style and stigma lobes white to light green.

inner tepals green- or orange-bronze, spatulate, 17–20 mm, apiculate;

filaments dark green-bronze;

anthers yellow;

style whitish basally to light orange apically;

stigma lobes very pale green.

Fruits

tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus;

basal tubercles longest;

areoles 12–30.

green, becoming yellowish apically, sometimes tinged red to purplish at areoles, commonly sterile, then narrow, tuberculate, to 2.5 cm, fertile ones stipitate, obconic, 20–50 × 15–35 mm, fleshy, becoming smooth, spineless;

umbilicus 3–4 mm deep;

areoles 15–17(–34).

Seeds

pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions;

girdle smooth.

pale yellow, angularly circular to oblong, thick, 3–5 mm diam., lumpy;

girdle smooth, broad, not protruding.

2n

= 66.

Cylindropuntia acanthocarpa

Cylindropuntia arbuscula

Phenology Flowering spring (Apr–Jun).
Habitat Sonoran Desert flats, bajadas, desert scrub
Elevation 300-1000 m (1000-3300 ft)
Distribution
from FNA
AZ; CA; NV; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; Mexico (Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cylindropuntia arbuscula forms hybrids with C. leptocaulis in south-central Arizona; those hybrids have narrow, obscurely tuberculate stems and reddish fruits nearly the size of those of C. arbuscula and chromosome number of 2n = 55. Hybrids between C. arbuscula and C. spinosior [= C. ×neoarbuscula (Griffiths) F. M. Knuth] have large green fruits, which often split open, and on stem segments distal areoles that usually bear three or four spines to 2.5 cm. Hybrids between C. arbuscula and C. versicolor [= C. ×vivipara (Rose) F. M. Knuth] have large green fruits, which often split open, and distal stem segment areoles that bear one or two spines to 1.5 cm.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tubercles 2.5-4.5 cm, broad; stem segments gray-green; spines of adjacent areoles not or barely interlaced, 6-11(-14) per areole; fruits bearing few spines or sometimes spineless
var. thornbe
1. Tubercles (1-)1.5-2.5(-3) cm, broad to narrow; stem segments green to yellow-green; spines of adjacent areoles interlaced; fruits densely spiny, sometimes burlike
→ 2
2. Abaxial 5-7 spines deflexed, usually subparallel, whitish to light brown; tubercles broad, oval; shrubs with few, ascending branches
var. acantho
2. Abaxial (14-)18-30+ spines deflexed to divergent, spreading, pale yellow to red-brown; tubercles narrowly elliptic; shrubs with few to many, spreading branches
→ 3
3. Branches usually at acute angles; stem segments yellowish green, usually long, commonly 15-30 cm; spines 18-30+, yellow to tan, sheaths baggy; inner tepals yellow
var. colorad
3. Branches usually at obtuse angles; stem segments green to dark green, usually short, commonly 8 -20 cm; spines 14-18(-23) per areole, yellow or red-brown, sheaths tight; inner tepals yellow to bronze to brick red
var. major
Source FNA vol. 4. FNA vol. 4, p. 106.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Subordinate taxa
C. acanthocarpa var. acantho, C. acanthocarpa var. colorad, C. acanthocarpa var. major, C. acanthocarpa var. thornbe
Synonyms Opuntia acanthocarpa Opuntia arbuscula
Name authority (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) (Engelmann) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 123. (1935)
Web links