Cylindropuntia acanthocarpa |
Cylindropuntia arbuscula |
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buck-horn cholla |
Arizona pencil cholla, pencil cholla |
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Habit | Trees or shrubs, sparsely to densely branched, 1–2(–4) m. Stem segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm; tubercles prominent, narrow to broad, 1.5–4.5 cm; areoles elliptic to subcircular, 4–5.5 × 4–5 mm; wool white or yellowish to tan, aging gray-black. | Trees or shrubs, densely branched, 0.5–3 m. Stem segments green, sometimes purple tinged, 6–10.5 × 0.5–1.3 cm; tubercles narrowly elongate, appearing as wrinkles when dry, 1–2 cm; areoles circular, 2–3 mm in diam.; wool tan to brown, aging gray to black. | ||||||||||||
Spines | 6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles; major abaxial spines deflexed to divergent, subterete to flattened; major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm; sheaths white to grayish with yellow to golden tips, baggy to tight fitting. |
0–2(–3) per areole, sparsely distributed along stem, usually deflexed, pale yellow or red-brown aging black, stout, the longest 0.8–3.5(–5) cm; sheaths loose fitting, yellowish brown. |
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Glochids | in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm. |
in adaxial tuft and marginal, encircling areole, pale yellow. |
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Flowers | inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate; filaments red; anthers yellow; style and stigma lobes white to light green. |
inner tepals green- or orange-bronze, spatulate, 17–20 mm, apiculate; filaments dark green-bronze; anthers yellow; style whitish basally to light orange apically; stigma lobes very pale green. |
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Fruits | tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus; basal tubercles longest; areoles 12–30. |
green, becoming yellowish apically, sometimes tinged red to purplish at areoles, commonly sterile, then narrow, tuberculate, to 2.5 cm, fertile ones stipitate, obconic, 20–50 × 15–35 mm, fleshy, becoming smooth, spineless; umbilicus 3–4 mm deep; areoles 15–17(–34). |
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Seeds | pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions; girdle smooth. |
pale yellow, angularly circular to oblong, thick, 3–5 mm diam., lumpy; girdle smooth, broad, not protruding. |
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2n | = 66. |
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Cylindropuntia acanthocarpa |
Cylindropuntia arbuscula |
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Phenology | Flowering spring (Apr–Jun). | |||||||||||||
Habitat | Sonoran Desert flats, bajadas, desert scrub | |||||||||||||
Elevation | 300-1000 m (1000-3300 ft) | |||||||||||||
Distribution |
AZ; CA; NV; UT; n Mexico
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AZ; Mexico (Sonora)
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Discussion | Varieties 4 (4 in the flora). Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cylindropuntia arbuscula forms hybrids with C. leptocaulis in south-central Arizona; those hybrids have narrow, obscurely tuberculate stems and reddish fruits nearly the size of those of C. arbuscula and chromosome number of 2n = 55. Hybrids between C. arbuscula and C. spinosior [= C. ×neoarbuscula (Griffiths) F. M. Knuth] have large green fruits, which often split open, and on stem segments distal areoles that usually bear three or four spines to 2.5 cm. Hybrids between C. arbuscula and C. versicolor [= C. ×vivipara (Rose) F. M. Knuth] have large green fruits, which often split open, and distal stem segment areoles that bear one or two spines to 1.5 cm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4. | FNA vol. 4, p. 106. | ||||||||||||
Parent taxa | Cactaceae > subfam. Opuntioideae > Cylindropuntia | Cactaceae > subfam. Opuntioideae > Cylindropuntia | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | Opuntia acanthocarpa | Opuntia arbuscula | ||||||||||||
Name authority | (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) | (Engelmann) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 123. (1935) | ||||||||||||
Web links |