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buck-horn cholla

chain-fruit cholla, jumping cholla

Habit Trees or shrubs, sparsely to densely branched, 1–2(–4) m. Trees 1–3 m; trunk divaricately branching; crown many branched, spreading.
Stem

segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm;

tubercles prominent, narrow to broad, 1.5–4.5 cm;

areoles elliptic to subcircular, 4–5.5 × 4–5 mm;

wool white or yellowish to tan, aging gray-black.

segments whorled or subwhorled, gray-green, often drying blackish, ± spiny throughout, terminal ones easily dislodged, 6–16(–23) × 2–3.5 cm;

tubercles salient, broadly oval, 0.8–1.3(–1.9) cm;

areoles obdeltate, 5–7(–10) × 2.5–4 mm;

wool gold to tan, aging gray to black.

Spines

6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles;

major abaxial spines deflexed to divergent, subterete to flattened;

major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm;

sheaths white to grayish with yellow to golden tips, baggy to tight fitting.

0–12(–18) per areole, at most areoles to nearly absent, yellowish, sometimes also pale pinkish, aging brown, interlaced or not with spines of adjacent areoles;

abaxial spines erect to deflexed, spreading, flattened basally, the longest to 3.5 cm;

adaxial spines erect or spreading, terete to subterete, longest to 2.5 cm;

sheaths uniformly whitish, yellowish to golden, baggy.

Glochids

in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm.

in adaxial tuft, sometimes also scattered along areole margins, yellow, 1–3 mm.

Flowers

inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate;

filaments red;

anthers yellow;

style and stigma lobes white to light green.

inner tepals usually reflexed, pink to magenta, obovate to ligulate, 12–16 mm, apiculate emarginate;

filaments pale pink to magenta;

anthers white to cream;

style pinkish;

stigma lobes whitish to pale yellow.

Fruits

tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus;

basal tubercles longest;

areoles 12–30.

proliferating, forming long, branching, pendent chains, at maturity gray-green, often stipitate, obconic, fleshy, shallowly tuberculate, usually spineless;

basal fruits 32–55 × 23–45 mm;

terminal fruits 2–3.3 × 1.3–2.3 cm;

tubercles becoming obscure;

umbilicus to 8 mm deep;

areoles 18–35.

Seeds

pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions;

girdle smooth.

pale yellow to brownish, angular to very irregular in outline, warped, 1.9 × 1.5–3.5 mm, sides with 1–2 large depressions, hilum pointed;

girdle smooth.

Cylindropuntia acanthocarpa

Cylindropuntia fulgida

Distribution
from FNA
AZ; CA; NV; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Intermediates are known between the varieties, which are largely sympatric in northern portion of range of the species.

Cylindropuntia fulgida forms hybrids with C. spinosior (see 6. C. ×kelvinensis) and C. leptocaulis. Hybrids, which are rare in south-central Arizona, have stems of intermediate diameter, (0–)1–5 spines per areole, one spine much longer than others, and spineless, yellowing, and often reddish fruits in chains of four to six, or more.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tubercles 2.5-4.5 cm, broad; stem segments gray-green; spines of adjacent areoles not or barely interlaced, 6-11(-14) per areole; fruits bearing few spines or sometimes spineless
var. thornbe
1. Tubercles (1-)1.5-2.5(-3) cm, broad to narrow; stem segments green to yellow-green; spines of adjacent areoles interlaced; fruits densely spiny, sometimes burlike
→ 2
2. Abaxial 5-7 spines deflexed, usually subparallel, whitish to light brown; tubercles broad, oval; shrubs with few, ascending branches
var. acantho
2. Abaxial (14-)18-30+ spines deflexed to divergent, spreading, pale yellow to red-brown; tubercles narrowly elliptic; shrubs with few to many, spreading branches
→ 3
3. Branches usually at acute angles; stem segments yellowish green, usually long, commonly 15-30 cm; spines 18-30+, yellow to tan, sheaths baggy; inner tepals yellow
var. colorad
3. Branches usually at obtuse angles; stem segments green to dark green, usually short, commonly 8 -20 cm; spines 14-18(-23) per areole, yellow or red-brown, sheaths tight; inner tepals yellow to bronze to brick red
var. major
1. Spines of stems interlaced with spines of adjacent areoles, the longest usually 2.5-3 cm; sheaths baggy; stem segments appearing spiny from afar, obscuring strongly mammillate tubercles beneath.
var. fulgida
1. Spines of stems not or little interlaced with spines of adjacent areoles, the longest usually 1-2 cm; sheaths tightly fitting; stem segments appearing spineless or nearly so from afar, exposing strongly mammillate tubercles beneath
var. mamillata
Source FNA vol. 4. Treatment author: Donald J. Pinkava. FNA vol. 4. Treatment author: Donald J. Pinkava.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Subordinate taxa
C. acanthocarpa var. acantho, C. acanthocarpa var. colorad, C. acanthocarpa var. major, C. acanthocarpa var. thornbe
C. fulgida var. fulgida, C. fulgida var. mamillata
Synonyms Opuntia acanthocarpa Opuntia fulgida
Name authority (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) (Engelmann) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 126. (1935)
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