Cylindropuntia acanthocarpa |
Cylindropuntia echinocarpa |
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buck-horn cholla |
golden cholla, silver cholla, silver or golden cholla, Wiggins' cholla |
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Habit | Trees or shrubs, sparsely to densely branched, 1–2(–4) m. | Trees or shrubs, spreading, densely branched, 0.5–2 m. | ||||||||||||
Stem | segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm; tubercles prominent, narrow to broad, 1.5–4.5 cm; areoles elliptic to subcircular, 4–5.5 × 4–5 mm; wool white or yellowish to tan, aging gray-black. |
segments usually firmly attached, terminal segments sometimes easily detached, green to light gray-green, cylindric, 3–7.5(–12) × 1–2.5 cm; tubercles salient, short oval, 0.4–1.3(–1.5) cm, moderately broad; areoles broadly elliptic, 4–4.5 × 2–3.5 mm; wool whitish, yellow to tan, aging gray. |
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Spines | 6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles; major abaxial spines deflexed to divergent, subterete to flattened; major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm; sheaths white to grayish with yellow to golden tips, baggy to tight fitting. |
(6–)10–17(–22) per areole, accompanied by 0–5 short bristlelike spines at areole margins, at most areoles, most prominent in adaxial ones, much interlaced with those of adjacent areoles, obscuring stems, sometimes brushlike; major abaxial spines whitish, yellow to tan, usually subterete to flattened, spreading to deflexed, the longest 20–30(–45) mm; major adaxial spines erect or spreading, white, pale yellow, or tan to brown (rarely deep red-brown), terete, the longest 22–35(–50) mm; sheaths whitish with yellow to golden tips, baggy. |
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Glochids | in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm. |
conspicuous in broad adaxial crescent, yellow, 3–4(–5) mm. |
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Flowers | inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate; filaments red; anthers yellow; style and stigma lobes white to light green. |
inner tepals light green to yellow-green, sometimes suffused with maroon or rose, spatulate, 20–23 mm, emarginate-apiculate; filaments greenish white or yellow, sometimes suffused with bronze or rose, but not red; anthers yellow; style and stigma lobes whitish, cream to light green. |
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Fruits | tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus; basal tubercles longest; areoles 12–30. |
tan at maturity, obconic to subspheric, 13–23 × 11–20 mm, dry, tuberculate, densely spiny, sometimes burlike, with apical flange around deep umbilicus; basal tubercles not markedly longer than distal ones; areoles 16–24(–36). |
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Seeds | pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions; girdle smooth. |
pale yellow, angular to squarish in outline, warped, 4–6 × 3.5–5 mm, sides smooth, each with 2–4 large depressions; girdle smooth (rarely with very narrow ridge). |
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2n | = 22. |
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Cylindropuntia acanthocarpa |
Cylindropuntia echinocarpa |
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Phenology | Flowering spring (Mar–Jun). | |||||||||||||
Habitat | Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates | |||||||||||||
Elevation | 50-1700 m [160-5600 ft] | |||||||||||||
Distribution |
AZ; CA; NV; UT; n Mexico
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AZ; CA; NV; Mexico (Baja California, Sonora)
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Discussion | Varieties 4 (4 in the flora). Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A dwarf form, with narrow terminal stem segments bearing few spines per areole, and one spine longer than the others, occurs on the very arid flats along the lower Colorado River in California and Arizona and has been recognized as Opuntia wigginsii L. D. Benson; however, numerous intermediate to more robust forms make recognition of the dwarf form untenable. Although exceptions occur, plants to the north and west in the range tend to be more spiny and to bear yellow to yellow-green flowers. Cylindropuntia echinocarpa hybridizes with C. acanthocarpa [= C. ×deserta (Griffiths) Pinkava (D. J. Pinkava 1999)], C. whipplei, and C. munzii. Cylindropuntia ×deserta, found in western Arizona and southern California, has reddish filaments, short tubercles, variably sized spines/areole, and dry spiny fruits. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4. | FNA vol. 4, p. 115. | ||||||||||||
Parent taxa | ||||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | Opuntia acanthocarpa | Opuntia echinocarpa, Cactus echinocarpus, C. wigginsii, Opuntia wigginsii | ||||||||||||
Name authority | (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) | (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) | ||||||||||||
Web links |