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buck-horn cholla

golden cholla, silver cholla, silver or golden cholla, Wiggins' cholla

Habit Trees or shrubs, sparsely to densely branched, 1–2(–4) m. Trees or shrubs, spreading, densely branched, 0.5–2 m.
Stem

segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm;

tubercles prominent, narrow to broad, 1.5–4.5 cm;

areoles elliptic to subcircular, 4–5.5 × 4–5 mm;

wool white or yellowish to tan, aging gray-black.

segments usually firmly attached, terminal segments sometimes easily detached, green to light gray-green, cylindric, 3–7.5(–12) × 1–2.5 cm;

tubercles salient, short oval, 0.4–1.3(–1.5) cm, moderately broad;

areoles broadly elliptic, 4–4.5 × 2–3.5 mm;

wool whitish, yellow to tan, aging gray.

Spines

6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles;

major abaxial spines deflexed to divergent, subterete to flattened;

major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm;

sheaths white to grayish with yellow to golden tips, baggy to tight fitting.

(6–)10–17(–22) per areole, accompanied by 0–5 short bristlelike spines at areole margins, at most areoles, most prominent in adaxial ones, much interlaced with those of adjacent areoles, obscuring stems, sometimes brushlike;

major abaxial spines whitish, yellow to tan, usually subterete to flattened, spreading to deflexed, the longest 20–30(–45) mm;

major adaxial spines erect or spreading, white, pale yellow, or tan to brown (rarely deep red-brown), terete, the longest 22–35(–50) mm;

sheaths whitish with yellow to golden tips, baggy.

Glochids

in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm.

conspicuous in broad adaxial crescent, yellow, 3–4(–5) mm.

Flowers

inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate;

filaments red;

anthers yellow;

style and stigma lobes white to light green.

inner tepals light green to yellow-green, sometimes suffused with maroon or rose, spatulate, 20–23 mm, emarginate-apiculate;

filaments greenish white or yellow, sometimes suffused with bronze or rose, but not red;

anthers yellow;

style and stigma lobes whitish, cream to light green.

Fruits

tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus;

basal tubercles longest;

areoles 12–30.

tan at maturity, obconic to subspheric, 13–23 × 11–20 mm, dry, tuberculate, densely spiny, sometimes burlike, with apical flange around deep umbilicus;

basal tubercles not markedly longer than distal ones;

areoles 16–24(–36).

Seeds

pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions;

girdle smooth.

pale yellow, angular to squarish in outline, warped, 4–6 × 3.5–5 mm, sides smooth, each with 2–4 large depressions;

girdle smooth (rarely with very narrow ridge).

2n

= 22.

Cylindropuntia acanthocarpa

Cylindropuntia echinocarpa

Phenology Flowering spring (Mar–Jun).
Habitat Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates
Elevation 50-1700 m [160-5600 ft]
Distribution
from FNA
AZ; CA; NV; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

A dwarf form, with narrow terminal stem segments bearing few spines per areole, and one spine longer than the others, occurs on the very arid flats along the lower Colorado River in California and Arizona and has been recognized as Opuntia wigginsii L. D. Benson; however, numerous intermediate to more robust forms make recognition of the dwarf form untenable. Although exceptions occur, plants to the north and west in the range tend to be more spiny and to bear yellow to yellow-green flowers.

Cylindropuntia echinocarpa hybridizes with C. acanthocarpa [= C. ×deserta (Griffiths) Pinkava (D. J. Pinkava 1999)], C. whipplei, and C. munzii. Cylindropuntia ×deserta, found in western Arizona and southern California, has reddish filaments, short tubercles, variably sized spines/areole, and dry spiny fruits.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tubercles 2.5-4.5 cm, broad; stem segments gray-green; spines of adjacent areoles not or barely interlaced, 6-11(-14) per areole; fruits bearing few spines or sometimes spineless
var. thornbe
1. Tubercles (1-)1.5-2.5(-3) cm, broad to narrow; stem segments green to yellow-green; spines of adjacent areoles interlaced; fruits densely spiny, sometimes burlike
→ 2
2. Abaxial 5-7 spines deflexed, usually subparallel, whitish to light brown; tubercles broad, oval; shrubs with few, ascending branches
var. acantho
2. Abaxial (14-)18-30+ spines deflexed to divergent, spreading, pale yellow to red-brown; tubercles narrowly elliptic; shrubs with few to many, spreading branches
→ 3
3. Branches usually at acute angles; stem segments yellowish green, usually long, commonly 15-30 cm; spines 18-30+, yellow to tan, sheaths baggy; inner tepals yellow
var. colorad
3. Branches usually at obtuse angles; stem segments green to dark green, usually short, commonly 8 -20 cm; spines 14-18(-23) per areole, yellow or red-brown, sheaths tight; inner tepals yellow to bronze to brick red
var. major
Source FNA vol. 4. Treatment author: Donald J. Pinkava. FNA vol. 4, p. 115. Treatment author: Donald J. Pinkava.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Subordinate taxa
C. acanthocarpa var. acantho, C. acanthocarpa var. colorad, C. acanthocarpa var. major, C. acanthocarpa var. thornbe
Synonyms Opuntia acanthocarpa Opuntia echinocarpa, Cactus echinocarpus, C. wigginsii, Opuntia wigginsii
Name authority (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935)
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