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buck-horn cholla

California cholla, Snake cholla

Habit Trees or shrubs, sparsely to densely branched, 1–2(–4) m. Shrubs, decumbent to erect, essentially trunkless, 0.3–2.5 m.
Stem

segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm;

tubercles prominent, narrow to broad, 1.5–4.5 cm;

areoles elliptic to subcircular, 4–5.5 × 4–5 mm;

wool white or yellowish to tan, aging gray-black.

segments green to purplish green, 6–30 × 1.5–2.5 cm;

tubercles prominent, oval to elongate-elliptic, 0.7–3 cm, sometimes appearing as ribs;

areoles elliptic, 4.5–5 × 3–3.5 mm;

wool tan, aging dark gray.

Spines

6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles;

major abaxial spines deflexed to divergent, subterete to flattened;

major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm;

sheaths white to grayish with yellow to golden tips, baggy to tight fitting.

(0–)7–13(–20) per areole, at most areoles, rarely obscuring stems, yellow to orange-brown with yellow tips, aging dark gray, subequal or 1–6(–7) much longer to 3 cm;

sheaths whitish or gold to brown, moderately loose.

Glochids

in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm.

yellow-tan to red-brown in dense adaxial crescent and scattered, stouter ones at basal margins.

Flowers

inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate;

filaments red;

anthers yellow;

style and stigma lobes white to light green.

inner tepals yellow to greenish yellow;

outer ones tipped red, 15–30 mm;

filaments green or yellow;

anthers yellow;

style light green or cream-white (rarely pinkish) apically;

stigma lobes cream-white to yellow.

Fruits

tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus;

basal tubercles longest;

areoles 12–30.

green to yellowish, drying gray-tan, short top-shaped to subspheric, 15–32 × 13–26 mm, leathery-dry, tuberculate, few spined or spineless;

proximal tubercles longest;

umbilicus deep;

rim slightly flanged;

areoles 13–30.

Seeds

pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions;

girdle smooth.

tan, squarish to angular in outline, warped, (4–)5–7 × 4–5 mm, sides with 2–4 deep depressions;

girdle smooth.

Cylindropuntia acanthocarpa

Cylindropuntia californica

Distribution
from FNA
AZ; CA; NV; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tubercles 2.5-4.5 cm, broad; stem segments gray-green; spines of adjacent areoles not or barely interlaced, 6-11(-14) per areole; fruits bearing few spines or sometimes spineless
var. thornbe
1. Tubercles (1-)1.5-2.5(-3) cm, broad to narrow; stem segments green to yellow-green; spines of adjacent areoles interlaced; fruits densely spiny, sometimes burlike
→ 2
2. Abaxial 5-7 spines deflexed, usually subparallel, whitish to light brown; tubercles broad, oval; shrubs with few, ascending branches
var. acantho
2. Abaxial (14-)18-30+ spines deflexed to divergent, spreading, pale yellow to red-brown; tubercles narrowly elliptic; shrubs with few to many, spreading branches
→ 3
3. Branches usually at acute angles; stem segments yellowish green, usually long, commonly 15-30 cm; spines 18-30+, yellow to tan, sheaths baggy; inner tepals yellow
var. colorad
3. Branches usually at obtuse angles; stem segments green to dark green, usually short, commonly 8 -20 cm; spines 14-18(-23) per areole, yellow or red-brown, sheaths tight; inner tepals yellow to bronze to brick red
var. major
1. Shrubs, suberect to decumbent; stem tubercles oval, shorter, usually 0.7-1.5 cm; spines/areole usually subequal
var. californ
1. Shrubs, suberect to erect; stem tubercles elliptic, longer, usually 1.5-3 cm; spines/areole commonly with 1(-3) central spines much longer
var. parkeri
Source FNA vol. 4. Treatment author: Donald J. Pinkava. FNA vol. 4, p. 113. Treatment author: Donald J. Pinkava.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Subordinate taxa
C. acanthocarpa var. acantho, C. acanthocarpa var. colorad, C. acanthocarpa var. major, C. acanthocarpa var. thornbe
C. californica var. californ, C. californica var. parkeri
Synonyms Opuntia acanthocarpa Cereus californicus, Opuntia californica
Name authority (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) (Torrey & A. Gray) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. (1935)
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