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buck-horn cholla

coyonostyle, tree cholla

Habit Trees or shrubs, sparsely to densely branched, 1–2(–4) m. Stem segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm; tubercles prominent, narrow to broad, 1.5–4.5 cm; areoles elliptic to subcircular, 4–5.5 × 4–5 mm; wool white or yellowish to tan, aging gray-black. Trees, with short trunks, or large shrubs, widely branching, (1–)1.5–2.5(–5) m. Stem segments firmly attached, whorled or subwhorled, gray-green, cylindric to weakly clavate, 10–40 × 1.5–4 cm; tubercles very prominent, widely spaced, (1.5–)2–5 cm; areoles elliptic, 5–8× 3–4 mm; wool yellow to tan, aging gray to black.
Spines

6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles;

major abaxial spines deflexed to divergent, subterete to flattened;

major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm;

sheaths white to grayish with yellow to golden tips, baggy to tight fitting.

(5–)8–15(–30) per areole or spineless to nearly spineless, usually at most areoles, not obscuring stems, spreading, straight or slightly curved, silver to yellow to usually ± red or tan to brown, stout, terete or sometimes flattened basally, 8–30(–40) mm subequal;

sheaths silver to yellow to usually tan to dirty white, yellow tipped.

Glochids

in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm.

in dense adaxial tuft, pale yellow, 0.5–3 mm.

Flowers

inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate;

filaments red;

anthers yellow;

style and stigma lobes white to light green.

inner tepals dark pink to magenta to red-magenta, obovate, apiculate, 15–35 mm;

filaments green basally to pink to magenta distally;

anthers yellow;

style light green basally or pink to red-magenta apically;

stigma lobes green or cream.

Fruits

tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus;

basal tubercles longest;

areoles 12–30.

not proliferating, yellow, obovoid, 24–45 × 20–40 mm, fleshy, tuberculate, spineless;

areoles 18–30;

tubercles nearly equal in length or longer in proximal portion of fruit, prominent (occasionally smooth at maturity);

umbilicus 7–14 mm deep.

Seeds

pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions;

girdle smooth.

yellow-tan, subcircular to angled, warped, 2.5–4 mm diam., sides smooth to slightly lumpy;

girdle usually narrow, not protruding.

Cylindropuntia acanthocarpa

Cylindropuntia imbricata

Distribution
from FNA
AZ; CA; NV; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; KS; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2+ (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tubercles 2.5-4.5 cm, broad; stem segments gray-green; spines of adjacent areoles not or barely interlaced, 6-11(-14) per areole; fruits bearing few spines or sometimes spineless
var. thornbe
1. Tubercles (1-)1.5-2.5(-3) cm, broad to narrow; stem segments green to yellow-green; spines of adjacent areoles interlaced; fruits densely spiny, sometimes burlike
→ 2
2. Abaxial 5-7 spines deflexed, usually subparallel, whitish to light brown; tubercles broad, oval; shrubs with few, ascending branches
var. acantho
2. Abaxial (14-)18-30+ spines deflexed to divergent, spreading, pale yellow to red-brown; tubercles narrowly elliptic; shrubs with few to many, spreading branches
→ 3
3. Branches usually at acute angles; stem segments yellowish green, usually long, commonly 15-30 cm; spines 18-30+, yellow to tan, sheaths baggy; inner tepals yellow
var. colorad
3. Branches usually at obtuse angles; stem segments green to dark green, usually short, commonly 8 -20 cm; spines 14-18(-23) per areole, yellow or red-brown, sheaths tight; inner tepals yellow to bronze to brick red
var. major
1. Plants trees 3-5 m; stem segments 12-40 cm; tubercles widely spaced; spines, and sheaths when present, usually tan to dirty white or ± yellow
var. imbricata
1. Plants erect shrubs to 1.2 m; stem segments 10-20 cm; tubercles narrowly spaced; spines and sheaths silvery
var. argentea
Source FNA vol. 4. FNA vol. 4, p. 112.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Subordinate taxa
C. acanthocarpa var. acantho, C. acanthocarpa var. colorad, C. acanthocarpa var. major, C. acanthocarpa var. thornbe
C. imbricata var. argentea, C. imbricata var. imbricata
Synonyms Opuntia acanthocarpa family cactaceae Cereus imbricatusH, Opuntia imbricata
Name authority (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) (Haworth) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. (1935)
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