Johanneshowellia puberula |
Polygonaceae subfam. eriogonoideae |
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downy buckwheat, red creek buckwheat, red creek Howell's-buckwheat |
wild buckwheat |
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Habit | Plants weakly erect, grayish to greenish, 0.5–3 × 1–2 dm, mostly densely silky-puberulent. | Shrubs, subshrubs, or herbs, sometimes nearly arborescent (Eriogonum), perennial, biennial, or annual, homophyllous, polycarpic (rarely monocarpic in Eriogonum); taproot solid or, rarely, chambered (Eriogonum), slender to stout. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | 0.3–0.8 dm. |
prostrate or decumbent to spreading or erect, sometimes scapose, rarely absent (Eriogonum), without recurved spines, glabrous or pubescent, sometimes glandular; nodes not swollen; tendrils absent; caudex stems tightly compact to spreading and at or just below the soil surface or spreading to erect and above the soil surface, woody; aerial flowering stems decumbent to spreading or erect, arising at nodes of caudex branches, at distal nodes of aerial branches, or directly from the root, slender to stout and solid or slightly to distinctly fistulose, rarely disarticulating in ringlike segments (Eriogonum). |
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Leaves | blades obovate to rounded, 0.5–1.5 × 0.5–1.5 cm. |
deciduous (persistent in some shrubby and matted Eriogonum species), basal or basal and cauline, rarely only cauline, rosulate, alternate, or infrequently opposite (Goodmania) or in whorls of 3 (Gilmania); stipules absent (possibly vestigial in some perennial species of Chorizanthe); petiole present, sometimes indistinct, not articulate or with extrafloral nectaries; blade simple, rarely lobed (Pterostegia), rarely awn-tipped (Goodmania). |
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Inflorescences | spreading, open, 5–25 cm; bracts linear to narrowly lanceolate, 2–5(–9) × 1–2(–2.5) mm, somewhat foliaceous, puberulent. |
terminal or terminal and axillary, cymose and dichotomously or trichotomously branched, or racemose, simple or compound umbellate, or capitate; bracts usually connate proximally, leaflike or scalelike, entire apically, sometimes awn-tipped, glabrous or pubescent. |
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Peduncles | absent or erect to deflexed relative to inflorescence branch, sometimes reflexed, straight or curved. |
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Flowers | perianth white to pale yellow, becoming rose or red in fruit, 1.5–2 mm, minutely pustulose basally and along midribs; tepals slightly dimorphic, those of outer whorl narrowly ovate, slightly auriculate basally with undulate-crisped margins in fruit, those of inner whorl narrowly oblanceolate, often shorter than those of outer whorl; filaments 0.8–1.2 mm. |
(1–)2–30(–100) per involucral structure, occasionally with stipelike base distal to articulations (Eriogonum); perianth accrescent in fruit, mostly white to red, yellow, light green, greenish white, maroon, or purple, urceolate to campanulate, occasionally glandular or pustulose abaxially, nearly always minutely glandular along midvein adaxially, glabrous or pubescent; tepals (5–)6, in 2 whorls of 3, connate proximally, typically not forming tube (except Chorizanthe, Lastarriaea, Mucronea, Pterostegia), petaloid or, rarely, coriaceous (Lastarriaea), monomorphic or dimorphic, entire, emarginate, or lobed to laciniate apically, rarely awn-tipped (Lastarriaea) or apiculate (Eriogonum); nectary a disk at base of ovary; stamens 3, 6, or 9 (variously 3–9 in Chorizanthe, Mucronea); staminodes absent; filaments usually distinct, occasionally forming staminal tube (Chorizanthe); pistils 3-carpellate, homostylous; ovary 1-locular; ovule 1, orthotropous, placentation basal; styles 3, distinct; stigmas capitate. |
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Achenes | 1–1.5 mm. |
brown to black or maroon, homocarpic, winged or unwinged, 3-gonous, less often lenticular or globose-lenticular to globose. |
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Seeds | embryo straight or curved. |
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Involucral | bracts oblanceolate, 0.5–1.2 × 0.1–0.3 mm, villous abaxially, outermost one 2- or 3-lobed apically. |
structures tubular (involucre) or consisting of a series of individual bractlike lobes (involucral bracts) arranged in whorls or spirals, rarely absent (Gilmania), awns present or absent; involucre cylindric, prismatic, turbinate, campanulate, urceolate, or funnelform with 3–8(–36) usually erect teeth or 4–12 spreading to reflexed lobes (teeth and lobes are distal portions of proximally connate involucral bracts); involucral bracts in 1–3 whorls, rarely in spirals (Johanneshowellia), free or connate only at base, linear to oblanceolate or ovate. |
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Johanneshowellia puberula |
Polygonaceae subfam. eriogonoideae |
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Phenology | Flowering May–Sep. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Sandy flats and slopes, saltbush and sagebrush communities, pinyon-juniper woodlands | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | (500-)800-2800 m ((1600-)2600-9200 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; NV; UT |
Mainly temperate regions of w North America (Alaska to Mexico); uncommon in South America (Argentina and Chile) and e North America (WVa s to c Fla, e to Mo, Okla, and Tex) |
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Discussion | Johanneshowellia puberula is infrequent to occasionally locally common. Populations are widely scattered from the Cottonwood Mountains of Inyo County, California, across northern Clark, southern Eureka, Lincoln, Nye, and White Pine counties of Nevada, into Beaver, Iron, Millard, and Washington counties in Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 20, species ca. 325 (19 genera, 281 species in the flora). Detailed habitat, elevation, and distribution data for the eriogonoid genera are maintained by the author and available on the Web at: “Eriogonoideae (Polygonaceae) of North America north of Mexico” (http://www.life.umd.edu/emeritus/reveal/pbio/eriog/key.html). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 443. | FNA vol. 5, p. 218. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Eriogonum puberulum, Eriogonum puberulum var. venosum | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (S. Watson) Reveal: Brittonia 56: 302. (2004) | Arnott: in M. Napier, Encycl. Brit. ed. 7, 5: 126. (1832) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |