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Howell's-buckwheat

Habit Herbs, annual; taproot slender.
Stems

arising directly from the root, weakly erect to spreading, solid, not fistulose or disarticulating into ringlike segments, silky-puberulent.

Leaves

usually quickly deciduous, basal, rosulate;

petiole present;

blade obovate to rounded or somewhat reniform, margins entire.

Inflorescences

terminal, cymose;

branches dichotomous or trichotomous at proximal node, otherwise dichotomous, not brittle or disarticulating into segments, round, silky-puberulent;

bracts 3, connate proximally, triangular or linear to narrowly lanceolate, scalelike or somewhat leaflike, not awned, glabrous to puberulent.

Peduncles

absent.

Flowers

4–7 per involucral cluster;

perianth white or pale yellowish to rose or red, broadly campanulate when open, narrowly urceolate when closed, smooth or minutely pustulose abaxially, glabrous;

tepals 6, connate 1/4 their length, monomorphic or slightly dimorphic, entire apically;

stamens 9;

filaments basally adnate, glabrous;

anthers white to pale pink, oblong.

Achenes

included, light brown, not winged, 3-gonous, glabrous.

Seeds

embryo curved.

Involucral

bracts obscure, in tight spiral of (3–)4(–7) distinct, oblanceolate to obovate lobes, bractlike, not awned.

Johanneshowellia

Distribution
from USDA
w United States
[BONAP county map]
Discussion

Species 2 (2 in the flora).

With recognition of Johanneshowellia, all members of Eriogonum have their involucral bracts fused into a distinct, turbinate to campanulate or hemispheric, tubular structure. In Johanneshowellia, the four to seven, distinct, involucral bracts are arranged in a tight spiral. Each 0.5–1.2 mm, awnless, entire to deeply lobed bract subtends a flower-bearing pedicel. These involucral bracts are themselves usually obscured by inflorescence bracts that subtend each node. As a result, care must be taken to observe the true nature of the involucral complex. The genus is allied with Eriogonum subg. Oregonium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perianths minutely pustulose basally and along midribs, 1.5-2 mm in fruit; outer tepals narrowly ovate, slightly auriculate basally with slightly undulate-crisped margins in fruit; plants grayish to greenish, mostly densely silky-puberulent
J. puberula
1. Perianths smooth, 2-2.5 mm in fruit; tepals lanceolate, not auriculate basally, margins entire; plants reddish, thinly silky-puberulent
J. crateriorum
Source FNA vol. 5, p. 443. Author: James L. Reveal.
Parent taxa Polygonaceae > subfam. Eriogonoideae
Subordinate taxa
J. crateriorum, J. puberula
Synonyms Eriogonum unranked Puberula
Name authority Reveal: Brittonia 56: 299. (2004)
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