FNA: Diplacus calycinus

	Diplacus calycinus
	Kaweah River bush monkeyflower, rock bush monkeyflower
Habit	Subshrubs.
Stems	erect, 150–1500 mm, glandular-pubescent to viscid-villous.
Leaves	cauline, relatively even-sized; petiole absent; blade elliptic-lanceolate or lanceolate to narrowly lanceolate or linear-lanceolate, sometimes narrowly oblong, 20–75(–100) × 4–20(–28) mm, margins entire or shallowly crenate, plane or revolute, apex acute to obtuse, abaxial surfaces moderately villous, hairs unbranched, vitreous, adaxial glabrous.
Pedicels	3–5 mm in fruit.
Flowers	2 per node, chasmogamous.
Styles	minutely glandular.
Corollas	usually pale yellow or cream to yellow, not spotted or striped, palate ridges yellow to golden yellow, tube-throat 35–42 mm, limb 20–30 mm diam., bilabiate, lobes oblong, apex of adaxial 2 each shallowly, asymmetrically incised.
Calyces	not inflated in fruit, 28–40 mm, densely glandular-pubescent to short glandular-villous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green.
Capsules	25–35 mm.
Anthers	exserted, glabrous.
Stigmas	exserted, lobes equal.
2n	= 20.

	Diplacus calycinus
Phenology	Flowering Apr–Jul.
Habitat	Granite outcrops, boulders, rocky gullies.
Elevation	(300–)700–2200 m. ((1000–)2300–7200 ft.)
Distribution	CA [WildflowerSearch map] [BONAP county map]
Discussion	Although first described as a separate species, Diplacus calycinus has more recently been treated at subspecific or varietal rank (A. L. Grant 1924; F. W. Pennell 1951; P. A. Munz and D. D. Keck 1973). D. M. Thompson (2005) included both D. calycinus and D. longiflorus within his concept of Mimulus aurantiacus var. pubescens (Torrey) D. M. Thompson. He did not reference the study of sect. Diplacus by M. C. Tulig (2000), but results from the Tulig morphometric analyses indicated that D. calycinus is distinct from D. longiflorus, especially in corolla length, corolla tube length, and style length. The type of Diplacus calycinus is from Tulare County, and the concept of the species is perhaps best restricted to the Sierran population system in Fresno, Kern, and Tulare counties, disjunct from D. longiflorus, which occurs primarily in coastal counties. The Sierran system is characterized by distinct abaxial leaf vestiture; the hairs are unbranched, broad, and vitreous, compared to the branched, thinner, and dull hairs of D. longiflorus. Plants of D. calycinus parapatric with D. longiflorus also show a tendency toward the characteristic vestiture and also have lighter-colored (but more variable in color) corollas with narrower but slightly shorter tubes. Intergradation between D. calycinus and D. longiflorus occurs in the region connecting the San Bernardino and San Gabriel mountains in San Bernardino County. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 449.
Parent taxa	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	D. longiflorus var. calycinus, Mimulus longiflorus subsp. calycinus, M. longiflorus var. calycinus
Name authority	Eastwood: Bot. Gaz. 41: 287. (1906)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus calycinus

Kaweah River bush monkeyflower, rock bush monkeyflower

Habit

Subshrubs.

Stems

erect, 150–1500 mm, glandular-pubescent to viscid-villous.

Leaves

cauline, relatively even-sized;

petiole absent;

blade elliptic-lanceolate or lanceolate to narrowly lanceolate or linear-lanceolate, sometimes narrowly oblong, 20–75(–100) × 4–20(–28) mm, margins entire or shallowly crenate, plane or revolute, apex acute to obtuse, abaxial surfaces moderately villous, hairs unbranched, vitreous, adaxial glabrous.

Pedicels

3–5 mm in fruit.

Flowers

2 per node, chasmogamous.

Styles

minutely glandular.

Corollas

usually pale yellow or cream to yellow, not spotted or striped, palate ridges yellow to golden yellow, tube-throat 35–42 mm, limb 20–30 mm diam., bilabiate, lobes oblong, apex of adaxial 2 each shallowly, asymmetrically incised.

Calyces

not inflated in fruit, 28–40 mm, densely glandular-pubescent to short glandular-villous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green.

Capsules

25–35 mm.

Anthers

exserted, glabrous.

Stigmas

exserted, lobes equal.

= 20.

Diplacus calycinus

Phenology

Flowering Apr–Jul.

Habitat

Granite outcrops, boulders, rocky gullies.

Elevation

(300–)700–2200 m. ((1000–)2300–7200 ft.)

Distribution

[WildflowerSearch map]

[BONAP county map]

Discussion

Although first described as a separate species, Diplacus calycinus has more recently been treated at subspecific or varietal rank (A. L. Grant 1924; F. W. Pennell 1951; P. A. Munz and D. D. Keck 1973). D. M. Thompson (2005) included both D. calycinus and D. longiflorus within his concept of Mimulus aurantiacus var. pubescens (Torrey) D. M. Thompson. He did not reference the study of sect. Diplacus by M. C. Tulig (2000), but results from the Tulig morphometric analyses indicated that D. calycinus is distinct from D. longiflorus, especially in corolla length, corolla tube length, and style length.

The type of Diplacus calycinus is from Tulare County, and the concept of the species is perhaps best restricted to the Sierran population system in Fresno, Kern, and Tulare counties, disjunct from D. longiflorus, which occurs primarily in coastal counties. The Sierran system is characterized by distinct abaxial leaf vestiture; the hairs are unbranched, broad, and vitreous, compared to the branched, thinner, and dull hairs of D. longiflorus. Plants of D. calycinus parapatric with D. longiflorus also show a tendency toward the characteristic vestiture and also have lighter-colored (but more variable in color) corollas with narrower but slightly shorter tubes. Intergradation between D. calycinus and D. longiflorus occurs in the region connecting the San Bernardino and San Gabriel mountains in San Bernardino County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 449.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

D. longiflorus var. calycinus, Mimulus longiflorus subsp. calycinus, M. longiflorus var. calycinus

Name authority

Eastwood: Bot. Gaz. 41: 287. (1906)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Diplacus calycinus

Diplacus calycinus