FNA: Diplacus calycinus vs. Diplacus traskiae

	Diplacus calycinus	Diplacus traskiae
	Kaweah River bush monkeyflower, rock bush monkeyflower	Santa Catalina Island monkeyflower
Habit	Subshrubs.	Herbs, annual.
Stems	erect, 150–1500 mm, glandular-pubescent to viscid-villous.	erect, 80–120 mm, glandular-pubescent.
Leaves	cauline, relatively even-sized; petiole absent; blade elliptic-lanceolate or lanceolate to narrowly lanceolate or linear-lanceolate, sometimes narrowly oblong, 20–75(–100) × 4–20(–28) mm, margins entire or shallowly crenate, plane or revolute, apex acute to obtuse, abaxial surfaces moderately villous, hairs unbranched, vitreous, adaxial glabrous.	cauline, nearly even-sized or enlarging distally; petiole absent or with petiole-like extension; blade ovate to obovate, 12–41 × 5–21 mm, margins entire or crenate, plane, usually ciliate proximally, apex obtuse, surfaces glandular-pubescent.
Pedicels	3–5 mm in fruit.	3–5 mm in fruit.
Flowers	2 per node, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.
Styles	minutely glandular.	puberulent distally.
Corollas	usually pale yellow or cream to yellow, not spotted or striped, palate ridges yellow to golden yellow, tube-throat 35–42 mm, limb 20–30 mm diam., bilabiate, lobes oblong, apex of adaxial 2 each shallowly, asymmetrically incised.	throat magenta, abaxial lip magenta, adaxial lip whitish, palate ridges not seen, tube-throat 20–23 mm, limb 4–5 mm diam., bilabiate, lobes of abaxial lip smaller than adaxial.
Calyces	not inflated in fruit, 28–40 mm, densely glandular-pubescent to short glandular-villous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green.	distinctly asymmetrically attached to pedicel, not inflated in fruit, 18–21 mm, glandular-pubescent, lobes subequal, apex acute, ribs usually green, intercostal areas whitish.
Capsules	25–35 mm.	not seen.
Anthers	exserted, glabrous.	(distal pair) nearly exserted, glabrous.
Stigmas	exserted, lobes equal.	exserted, lobes unequal, abaxial 6–8 times adaxial.
2n	= 20.

	Diplacus calycinus	Diplacus traskiae
Phenology	Flowering Apr–Jul.	Flowering Mar–Apr.
Habitat	Granite outcrops, boulders, rocky gullies.	Rocky, brushy slopes.
Elevation	(300–)700–2200 m. ((1000–)2300–7200 ft.)	0–100 m. (0–300 ft.)
Distribution	CA [WildflowerSearch map] [BONAP county map]	CA
Discussion	Although first described as a separate species, Diplacus calycinus has more recently been treated at subspecific or varietal rank (A. L. Grant 1924; F. W. Pennell 1951; P. A. Munz and D. D. Keck 1973). D. M. Thompson (2005) included both D. calycinus and D. longiflorus within his concept of Mimulus aurantiacus var. pubescens (Torrey) D. M. Thompson. He did not reference the study of sect. Diplacus by M. C. Tulig (2000), but results from the Tulig morphometric analyses indicated that D. calycinus is distinct from D. longiflorus, especially in corolla length, corolla tube length, and style length. The type of Diplacus calycinus is from Tulare County, and the concept of the species is perhaps best restricted to the Sierran population system in Fresno, Kern, and Tulare counties, disjunct from D. longiflorus, which occurs primarily in coastal counties. The Sierran system is characterized by distinct abaxial leaf vestiture; the hairs are unbranched, broad, and vitreous, compared to the branched, thinner, and dull hairs of D. longiflorus. Plants of D. calycinus parapatric with D. longiflorus also show a tendency toward the characteristic vestiture and also have lighter-colored (but more variable in color) corollas with narrower but slightly shorter tubes. Intergradation between D. calycinus and D. longiflorus occurs in the region connecting the San Bernardino and San Gabriel mountains in San Bernardino County. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus traskiae is known only from Santa Catalina Island. According to the California Native Plant Society Online Inventory of Rare and Endangered Plants, ed. 8 (http://www.cnps.org/cnps/rareplants/inventory/), it (as Mimulus traskiae) is possibly extirpated. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 449.	FNA vol. 17, p. 446.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	D. longiflorus var. calycinus, Mimulus longiflorus subsp. calycinus, M. longiflorus var. calycinus	Mimulus traskiae
Name authority	Eastwood: Bot. Gaz. 41: 287. (1906)	(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 32. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus traskiae

Kaweah River bush monkeyflower, rock bush monkeyflower

Santa Catalina Island monkeyflower

Habit

Subshrubs.

Herbs, annual.

Stems

erect, 150–1500 mm, glandular-pubescent to viscid-villous.

erect, 80–120 mm, glandular-pubescent.

Leaves

cauline, relatively even-sized;

petiole absent;

blade elliptic-lanceolate or lanceolate to narrowly lanceolate or linear-lanceolate, sometimes narrowly oblong, 20–75(–100) × 4–20(–28) mm, margins entire or shallowly crenate, plane or revolute, apex acute to obtuse, abaxial surfaces moderately villous, hairs unbranched, vitreous, adaxial glabrous.

cauline, nearly even-sized or enlarging distally;

petiole absent or with petiole-like extension;

blade ovate to obovate, 12–41 × 5–21 mm, margins entire or crenate, plane, usually ciliate proximally, apex obtuse, surfaces glandular-pubescent.

Pedicels

3–5 mm in fruit.

Flowers

2 per node, chasmogamous.

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

Styles

minutely glandular.

puberulent distally.

Corollas

usually pale yellow or cream to yellow, not spotted or striped, palate ridges yellow to golden yellow, tube-throat 35–42 mm, limb 20–30 mm diam., bilabiate, lobes oblong, apex of adaxial 2 each shallowly, asymmetrically incised.

throat magenta, abaxial lip magenta, adaxial lip whitish, palate ridges not seen, tube-throat 20–23 mm, limb 4–5 mm diam., bilabiate, lobes of abaxial lip smaller than adaxial.

Calyces

not inflated in fruit, 28–40 mm, densely glandular-pubescent to short glandular-villous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green.

distinctly asymmetrically attached to pedicel, not inflated in fruit, 18–21 mm, glandular-pubescent, lobes subequal, apex acute, ribs usually green, intercostal areas whitish.

Capsules

25–35 mm.

not seen.

Anthers

exserted, glabrous.

(distal pair) nearly exserted, glabrous.

Stigmas

exserted, lobes equal.

exserted, lobes unequal, abaxial 6–8 times adaxial.

= 20.

Diplacus calycinus

Diplacus traskiae

Phenology

Flowering Apr–Jul.

Flowering Mar–Apr.

Habitat

Granite outcrops, boulders, rocky gullies.

Rocky, brushy slopes.

Elevation

(300–)700–2200 m. ((1000–)2300–7200 ft.)

0–100 m. (0–300 ft.)

Distribution

[WildflowerSearch map]

[BONAP county map]

Discussion

Although first described as a separate species, Diplacus calycinus has more recently been treated at subspecific or varietal rank (A. L. Grant 1924; F. W. Pennell 1951; P. A. Munz and D. D. Keck 1973). D. M. Thompson (2005) included both D. calycinus and D. longiflorus within his concept of Mimulus aurantiacus var. pubescens (Torrey) D. M. Thompson. He did not reference the study of sect. Diplacus by M. C. Tulig (2000), but results from the Tulig morphometric analyses indicated that D. calycinus is distinct from D. longiflorus, especially in corolla length, corolla tube length, and style length.

The type of Diplacus calycinus is from Tulare County, and the concept of the species is perhaps best restricted to the Sierran population system in Fresno, Kern, and Tulare counties, disjunct from D. longiflorus, which occurs primarily in coastal counties. The Sierran system is characterized by distinct abaxial leaf vestiture; the hairs are unbranched, broad, and vitreous, compared to the branched, thinner, and dull hairs of D. longiflorus. Plants of D. calycinus parapatric with D. longiflorus also show a tendency toward the characteristic vestiture and also have lighter-colored (but more variable in color) corollas with narrower but slightly shorter tubes. Intergradation between D. calycinus and D. longiflorus occurs in the region connecting the San Bernardino and San Gabriel mountains in San Bernardino County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus traskiae is known only from Santa Catalina Island. According to the California Native Plant Society Online Inventory of Rare and Endangered Plants, ed. 8 (http://www.cnps.org/cnps/rareplants/inventory/), it (as Mimulus traskiae) is possibly extirpated.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 449.

FNA vol. 17, p. 446.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

D. longiflorus var. calycinus, Mimulus longiflorus subsp. calycinus, M. longiflorus var. calycinus

Mimulus traskiae

Name authority

Eastwood: Bot. Gaz. 41: 287. (1906)

(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 32. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Diplacus calycinus

Diplacus traskiae

Diplacus calycinus

Diplacus traskiae