Cheilanthes bonariensis |
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Bonaire lip fern, golden lipfern |
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Stems | short-creeping to compact, usually 4–8 mm diam.; scales bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, narrowly lanceolate, slightly contorted, strongly appressed, persistent. |
Leaves | clustered, 10–60 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
Blade | linear, pinnate-pinnatifid throughout, 1–4 cm wide; rachis rounded adaxially, lacking scales, with dense monomorphic pubescence. |
Ultimate segments | elongate-deltate to ovate, not especially beadlike, the largest 1–7 mm, abaxially densely tomentose, adaxially hirsute. |
Pinnae | articulate at swollen, hirsute nodes, basal pair slightly smaller than adjacent pair, ± equilateral, appearing hirsute adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
Sporangia | containing 32 spores. |
Costae | absent. |
n | = 2n = 90, apogamous. |
Cheilanthes bonariensis |
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Phenology | Sporulating summer–fall. |
Habitat | Rocky slopes and ledges, found on a variety of substrates though rarely observed on limestone |
Elevation | 1200–2400 m (3900–7900 ft) |
Distribution |
AZ; NM; TX; Mexico; Central America; South America; West Indies |
Discussion | Cheilanthes bonariensis has been assigned to Notholaena in past treatments. It is distantly related (at best) to the species here included in Notholaena, however, and we concur with R. M. Tryon and A. F. Tryon (1982) that it should be transferred to Cheilanthes. Chromosomal studies (G. J. Gastony and M. D. Windham 1989) suggest that C. bonariensis is an apogamous triploid that arose through autopolyploidy. Further investigation is necessary to determine whether 64-spored, sexually reproducing populations of C. bonariensis are still extant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes |
Sibling taxa | |
Synonyms | Acrostichum bonariense, Notholaena aurea |
Name authority | (Willdenow) Proctor: Bull. Inst. Jamaica, Sci. Ser. 5(1): 15. (1953) |
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