Cheilanthes bonariensis |
Cheilanthes aemula |
|
---|---|---|
Bonaire lip fern, golden lipfern |
rival lipfern, Texas lip fern |
|
Stems | short-creeping to compact, usually 4–8 mm diam.; scales bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, narrowly lanceolate, slightly contorted, strongly appressed, persistent. |
short-creeping, usually 4–7 mm diam.; scales uniformly brown or slightly darker at base, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. |
Leaves | clustered, 10–60 cm; vernation noncircinate. |
clustered, 10–50 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
black to dark brown, rounded adaxially. |
Blade | linear, pinnate-pinnatifid throughout, 1–4 cm wide; rachis rounded adaxially, lacking scales, with dense monomorphic pubescence. |
ovate-deltate, 3-pinnate to 3-pinnate-pinnatifid at base, 4–15 cm wide; rachis rounded adaxially, lacking scales, with dimorphic pubescence, abaxially sparsely hirsute, adaxially covered with tortuous, appressed hairs. |
Ultimate segments | elongate-deltate to ovate, not especially beadlike, the largest 1–7 mm, abaxially densely tomentose, adaxially hirsute. |
narrowly elliptic to elongate-deltate, not beadlike, the largest 3–6 mm, abaxially and adaxially sparsely hirsute to glabrescent. |
Pinnae | articulate at swollen, hirsute nodes, basal pair slightly smaller than adjacent pair, ± equilateral, appearing hirsute adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair slightly larger than adjacent pair, somewhat inequilateral, proximal basiscopic pinnules conspicuously enlarged, appearing glabrous or sparsely pubescent adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, slightly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
somewhat discontinuous, often concentrated on interrupted lateral lobes. |
Sporangia | containing 32 spores. |
containing 64 spores. |
Costae | absent. |
black adaxially for most of length; abaxial scales absent. |
n | = 2n = 90, apogamous. |
|
2n | = 58. |
|
Cheilanthes bonariensis |
Cheilanthes aemula |
|
Phenology | Sporulating summer–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes and ledges, found on a variety of substrates though rarely observed on limestone | Rocky slopes and ledges, apparently confined to limestone |
Elevation | 1200–2400 m (3900–7900 ft) | 100–500 m (300–1600 ft) |
Distribution |
AZ; NM; TX; Mexico; Central America; South America; West Indies |
TX; n Mexico |
Discussion | Cheilanthes bonariensis has been assigned to Notholaena in past treatments. It is distantly related (at best) to the species here included in Notholaena, however, and we concur with R. M. Tryon and A. F. Tryon (1982) that it should be transferred to Cheilanthes. Chromosomal studies (G. J. Gastony and M. D. Windham 1989) suggest that C. bonariensis is an apogamous triploid that arose through autopolyploidy. Further investigation is necessary to determine whether 64-spored, sexually reproducing populations of C. bonariensis are still extant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In addition to the characteristics mentioned in the key, Cheilanthes aemula is distinguished from North American populations of C. alabamensis by having 64 spores per sporangium rather than 32. In the flora, this species is known from about 10 localities in central and western Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | Acrostichum bonariense, Notholaena aurea | |
Name authority | (Willdenow) Proctor: Bull. Inst. Jamaica, Sci. Ser. 5(1): 15. (1953) | Maxon: Contr. U.S. Natl. Herb. 10: 495. (1908) |
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