Cheilanthes bonariensis |
Cheilanthes viscida |
|
---|---|---|
Bonaire lip fern, golden lipfern |
viscid lip fern |
|
Stems | short-creeping to compact, usually 4–8 mm diam.; scales bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, narrowly lanceolate, slightly contorted, strongly appressed, persistent. |
short-creeping, usually 4–8 mm diam.; scales uniformly brown, linear-subulate, strongly contorted, loosely appressed, persistent. |
Leaves | clustered, 10–60 cm; vernation noncircinate. |
clustered, 6–30 cm; vernation circinate. |
Petiole | dark brown, rounded adaxially. |
dark brown, flattened or slightly grooved distally on adaxial surface. |
Blade | linear, pinnate-pinnatifid throughout, 1–4 cm wide; rachis rounded adaxially, lacking scales, with dense monomorphic pubescence. |
narrowly oblong to linear, 3-pinnate-pinnatifid at base, 1–4 cm wide; rachis flattened or slightly grooved adaxially, lacking scales, with monomorphic pubescence. |
Ultimate segments | elongate-deltate to ovate, not especially beadlike, the largest 1–7 mm, abaxially densely tomentose, adaxially hirsute. |
oblong to lanceolate, not beadlike, the largest 3–4 mm, abaxially and adaxially glandular-pubescent with short, sticky, capitate glands. |
Pinnae | articulate at swollen, hirsute nodes, basal pair slightly smaller than adjacent pair, ± equilateral, appearing hirsute adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing glandular pubescent adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, weakly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
usually discontinuous, concentrated on apical and lateral lobes. |
Sporangia | containing 32 spores. |
containing 64 spores. |
Costae | absent. |
green adaxially for most of length; abaxial scales absent. |
n | = 2n = 90, apogamous. |
|
Cheilanthes bonariensis |
Cheilanthes viscida |
|
Phenology | Sporulating summer–fall. | Sporulating late spring–fall. |
Habitat | Rocky slopes and ledges, found on a variety of substrates though rarely observed on limestone | Cliffs and rocky slopes, usually on igneous substrates |
Elevation | 1200–2400 m (3900–7900 ft) | 200–1300 m (700–4300 ft) |
Distribution |
AZ; NM; TX; Mexico; Central America; South America; West Indies |
CA; Mexico in Baja California
|
Discussion | Cheilanthes bonariensis has been assigned to Notholaena in past treatments. It is distantly related (at best) to the species here included in Notholaena, however, and we concur with R. M. Tryon and A. F. Tryon (1982) that it should be transferred to Cheilanthes. Chromosomal studies (G. J. Gastony and M. D. Windham 1989) suggest that C. bonariensis is an apogamous triploid that arose through autopolyploidy. Further investigation is necessary to determine whether 64-spored, sexually reproducing populations of C. bonariensis are still extant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cheilanthes viscida is confined to a relatively small region in the deserts of California. Variations in spore size among populations suggest that the species may include more than one cytotype. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | Acrostichum bonariense, Notholaena aurea | |
Name authority | (Willdenow) Proctor: Bull. Inst. Jamaica, Sci. Ser. 5(1): 15. (1953) | Davenport: Bull. Torrey Bot. Club 6: 191. (1877) |
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