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Bonaire lip fern, golden lipfern

viscid lip fern

Stems

short-creeping to compact, usually 4–8 mm diam.;

scales bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, narrowly lanceolate, slightly contorted, strongly appressed, persistent.

short-creeping, usually 4–8 mm diam.;

scales uniformly brown, linear-subulate, strongly contorted, loosely appressed, persistent.

Leaves

clustered, 10–60 cm;

vernation noncircinate.

clustered, 6–30 cm;

vernation circinate.

Petiole

dark brown, rounded adaxially.

dark brown, flattened or slightly grooved distally on adaxial surface.

Blade

linear, pinnate-pinnatifid throughout, 1–4 cm wide;

rachis rounded adaxially, lacking scales, with dense monomorphic pubescence.

narrowly oblong to linear, 3-pinnate-pinnatifid at base, 1–4 cm wide;

rachis flattened or slightly grooved adaxially, lacking scales, with monomorphic pubescence.

Ultimate segments

elongate-deltate to ovate, not especially beadlike, the largest 1–7 mm, abaxially densely tomentose, adaxially hirsute.

oblong to lanceolate, not beadlike, the largest 3–4 mm, abaxially and adaxially glandular-pubescent with short, sticky, capitate glands.

Pinnae

articulate at swollen, hirsute nodes, basal pair slightly smaller than adjacent pair, ± equilateral, appearing hirsute adaxially.

not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing glandular pubescent adaxially.

False indusia

marginal, weakly differentiated, 0.05–0.25 mm wide.

marginal, weakly differentiated, 0.05–0.25 mm wide.

Sori

± continuous around segment margins.

usually discontinuous, concentrated on apical and lateral lobes.

Sporangia

containing 32 spores.

containing 64 spores.

Costae

absent.

green adaxially for most of length;

abaxial scales absent.

n

= 2n = 90, apogamous.

Cheilanthes bonariensis

Cheilanthes viscida

Phenology Sporulating summer–fall. Sporulating late spring–fall.
Habitat Rocky slopes and ledges, found on a variety of substrates though rarely observed on limestone Cliffs and rocky slopes, usually on igneous substrates
Elevation 1200–2400 m (3900–7900 ft) 200–1300 m (700–4300 ft)
Distribution
from FNA
AZ; NM; TX; Mexico; Central America; South America; West Indies
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
Discussion

Cheilanthes bonariensis has been assigned to Notholaena in past treatments. It is distantly related (at best) to the species here included in Notholaena, however, and we concur with R. M. Tryon and A. F. Tryon (1982) that it should be transferred to Cheilanthes. Chromosomal studies (G. J. Gastony and M. D. Windham 1989) suggest that C. bonariensis is an apogamous triploid that arose through autopolyploidy. Further investigation is necessary to determine whether 64-spored, sexually reproducing populations of C. bonariensis are still extant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cheilanthes viscida is confined to a relatively small region in the deserts of California. Variations in spore size among populations suggest that the species may include more than one cytotype.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Cheilanthes Pteridaceae > Cheilanthes
Sibling taxa
C. aemula, C. alabamensis, C. arizonica, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. wootonii, C. wrightii, C. yavapensis
Synonyms Acrostichum bonariense, Notholaena aurea
Name authority (Willdenow) Proctor: Bull. Inst. Jamaica, Sci. Ser. 5(1): 15. (1953) Davenport: Bull. Torrey Bot. Club 6: 191. (1877)
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