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dangling suncress, reflexed rock-cress, secund rock-cress

Habit Biennials or perennials; short-lived; sexual; caudex present or absent.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–7(–10.5) dm, densely pubescent proximally, trichomes short-stalked, 2–8-rayed, 0.1–0.2 mm, sparsely pubescent distally.

Basal leaves

blade oblanceolate, 2–7 mm wide, margins entire or shallowly dentate, sometimes ciliate at petiole base, surfaces densely pubescent, trichomes short-stalked, 5–10-rayed, 0.05–0.2 mm.

Cauline leaves

15–40, concealing stem proximally;

blade auricles 0.5–2.5 mm or, rarely, absent, surfaces of distalmost leaves pubescent.

Racemes

15–80(–140)-flowered, usually unbranched.

Flowers

pendent at anthesis;

sepals pubescent;

petals white to lavender, 4–8 × 0.8–2.2 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

reflexed, abruptly recurved at base, otherwise usually straight, 7–12(–18) mm, pubescent, trichomes appressed, branched.

Fruits

strongly reflexed, usually appressed to rachis, sometimes somewhat secund, straight, edges parallel, 3.5–9 cm × 0.9–1.8 mm;

valves glabrous or sparsely pubescent throughout;

ovules 60–116 per ovary;

style 0.5–2 mm.

Seeds

uniseriate, 1–1.4 × 0.9–1.2 mm;

wing continuous, to 0.1 mm wide.

2n

= 14.

Boechera retrofracta

Phenology Flowering Apr–Aug.
Habitat Rock outcrops, open hillsides, gravel bars and sandy banks in grassland, sagebrush, oak woodlands, and open conifer forests
Elevation 300-3300 m (1000-10800 ft)
Distribution
from FNA
AK; CA; ID; MI; MT; NV; OR; UT; WA; WY; AB; BC; MB; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996; N. H. Holmgren 2005b), B. retrofracta is easily distinguished from that species by having narrower (0.9–1.8 versus 2–2.5 mm), mostly non-secund fruits that are almost always appressed to rachises. The two taxa have allopatric distributions, with B. retrofracta found on the North American continent (mostly west of the Great Plains) and B. holboellii apparently confined to Greenland. Boechera retrofracta has formed hybrids with at least 12 other species. Besides differing in macromorphological characters, all those hybrids are distinct from B. retrofracta in the strict sense in having wider (20–30 versus 13–16 µm), spheroid pollen grains with asymmetric colpi.

Arabis kochii Blankinship is an illegitimate name, sometimes found in synonymy with Boechera retrofracta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 402.
Parent taxa Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis retrofracta, Arabis arcuata var. secunda, Arabis exilis, Arabis holboellii var. retrofracta, Arabis holboellii var. secunda, Arabis lignipes, Arabis retrofracta var. multicaulis, Arabis secunda, Arabis sparsiflora subsp. secunda, Arabis tenuis, B. exilis, B. holboellii var. secunda
Name authority (Graham) Á. Löve & D. Löve: Taxon 31: 125. (1982)
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