Viola nephrophylla |
Viola canadensis |
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kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle |
Canada violet, Canada white violet, Canadian violet, Canadian white violet, rugose violet |
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Habit | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. | Plants perennial, caulescent, not stoloniferous, 3–46(–60) cm, with branching rhizomes forming colonies or not. | ||||||||
Stems | 1–3(4), usually erect to ascending, glabrous or puberulent, from fleshy or subligneous rhizome. |
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Leaves | basal, 4–7, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–25 cm, usually glabrous; blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent. |
basal and cauline; basal: 1–5; stipules ± oblong, ovate, or lanceolate, margins entire, sometimes glandular, apex acuminate to cuspidate; petiole 1.1–23 cm, glabrous or puberulent; blade ovate to broadly ovate or ovate-reniform, 0.7–12.4 × 0.9–11.1(–12.3) cm, base cordate, subcordate, or truncate, margins crenate, crenulate, or serrulate, ciliate (sometimes only on proximal 1/2) or eciliate, apex acute to acuminate, surfaces glabrous or puberulent (often only on veins); cauline similar to basal except: stipules also deltate, margins also erose or laciniate, apex acute, long-acuminate to cuspidate, or ± truncate, occasionally 2- or 3-fid; petiole 0.1–6.9(–15.2) cm; blade ovate to deltate, 1.2–7.7 × 0.8–7.8 cm, base cordate to truncate, margins crenate or crenulate to ± serrulate. |
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Peduncles | 3–25 cm, usually glabrous. |
1–6.1 cm, glabrate to puberulent, sometimes glabrous below bracteoles. |
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Flowers | sepals ovate, margins usually eciliate, auricles 1–2 mm; petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on erect to ascending peduncles. |
sepals lanceolate, margins usually eciliate, auricles 0.5–1.3 mm; petals white adaxially, upper 2 and lower 3 tinged soft reddish violet abaxially, rarely white on both surfaces, all petals usually with yellow patch basally, lower 3 usually purple-veined, lateral 2 bearded, lowest 5.5–20 mm, spur white, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary or absent. |
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Capsules | ovoid, 5–10 mm, glabrous. |
ovoid to ellipsoid, 2.5–10 mm, sometimes muriculate, glabrous or puberulent. |
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Seeds | beige to brown or dark brown, 1.5–2.5 mm. |
brown to dark brown or purplish black, 1.5–2.5 mm. |
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2n | = 54. |
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Viola nephrophylla |
Viola canadensis |
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Phenology | Flowering Jan–Sep. | |||||||||
Habitat | Wet habitats in saturated soil in prairies, pastures, bogs, fens, sedge meadows, rocky shores of lakes and streams, limestone outcrops, gravelly calcareous stream beds | |||||||||
Elevation | 100–3000 m (300–9800 ft) | |||||||||
Distribution |
AR; AZ; CA; CO; CT; IA; ID; IL; KS; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; PA; SD; TX; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
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AK; AL; AR; AZ; CO; CT; GA; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OR; PA; RI; SC; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; NT; ON; QC; SK; YT
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Discussion | Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets. There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants. Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla. Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 139. | FNA vol. 6, p. 125. | ||||||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | V. arizonica, V. austiniae, V. cognata, V. crassula, V. galacifolia, V. lunellii, V. mccabeiana, V. nephrophylla var. arizonica, V. nephrophylla var. cognata, V. peramoena, V. pratincola, V. prionosepala, V. retusa, V. subjuncta, V. vagula | |||||||||
Name authority | Greene: Pittonia 3: 144. (1896) | Linnaeus: Sp. Pl. 2: 936. (1753) | ||||||||
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