Thelypodium wrightii |
Thelypodium howellii |
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Wright's stanleyella, Wright's thelypody |
Howell's thelypody |
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Habit | Biennials; slightly glaucous, glabrous. | Biennials or, rarely, perennials; (short-lived); glaucous, glabrous (except petioles). | ||||
Stems | branched distally, (6.5–)9.5–22.5(–28) dm. |
branched distally, 1–8(–9.3) dm. |
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Basal leaves | (and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm; blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate). |
petiole 0.5–2 cm, ciliate; blade often oblanceolate to spatulate, sometimes oblong or lanceolate, 2–10(–13.5) cm × 10–23(–40) mm, margins often lyrate to dentate, sometimes entire or repand. |
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Cauline leaves | petiolate; blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed. |
(ascending); sessile; blade often lanceolate to linear, sometimes oblong, (0.8–)1–9.7(–14.5) cm × 2–10(–30) mm, (base usually amplexicaul to strongly sagittate, rarely auriculate), margins entire. |
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Racemes | somewhat lax, considerably elongated in fruit. |
lax, elongated in fruit, (flower buds ovate to lanceolate). |
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Flowers | sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm; petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base); nectar glands confluent; filaments equal, (3–)3.5–6.5(–8.5) mm; anthers (1–)1.5–2.5(–3) mm, circinately coiled; gynophore 0.2–2(–5) mm. |
sepals erect, linear or lanceolate, (3.5–)5–8.5(–11.5) × (0.7–)1–2(–2.5) mm; petals lavender to purple, linear or oblanceolate to spatulate, (6–)9–16.5(–22) × 0.5–2.7(–3) mm, margins partially crisped, claw differentiated from blade, [slender, (2.5–)4–7(–11) mm, narrowest at base]; nectar glands confluent, subtending bases of stamens; filaments slightly tetradynamous, median pairs (3–)5.5–9(–12) mm, lateral pair (2.5–)4.5–8(–10) mm; anthers slightly exserted to included, linear, (1.5–)2–4(–5.5) mm, not circinately coiled; gynophore (stout), 0.5–1(–3.5) mm. |
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Fruiting pedicels | horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base. |
usually ascending, rarely divaricate-ascending, usually straight, rarely slightly incurved, stout, (2.5–)3.5–8(–14.5) mm, not flattened at base. |
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Fruits | horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds); ovules 76–128 per ovary; style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm. |
often ascending, sometimes erect or divaricate, slightly torulose, straight or slightly incurved (forming straight line with pedicel), terete, 1.5–6.5(–8.2) cm × 1–1.3(–2) mm; ovules 22–40 per ovary; style cylindrical or subclavate, (0.5–)1–2.7(–4.3) mm. |
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Seeds | 0.7–1.3(–1.5) × 0.5–0.8 mm. |
(1–)1.2–1.7(–2) × 0.5–0.7 mm. |
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Thelypodium wrightii |
Thelypodium howellii |
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Phenology | Flowering (Mar-)Jun–Oct. | |||||
Habitat | Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands | |||||
Elevation | 1200-2300 m (3900-7500 ft) | |||||
Distribution |
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
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CA; OR; WA
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Discussion | Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Streptanthus howellii (S. Watson) M. E. Jones is a later homonym of S. howellii S. Watson 1885. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 738. | FNA vol. 7, p. 732. | ||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae > Thelypodium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Stanleyella wrightii, T. wrightii subsp. oklahomensis | Cartiera howellii, Thelypodiopsis howellii | ||||
Name authority | A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) | S. Watson: Proc. Amer. Acad. Arts 21: 445. (1886) | ||||
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