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camphor tansy, dune tansy, eastern tansy, floccose tansy, Lake Huron tansy, tanaisie bipennée

tanaisie, tansy

Habit Perennials, 5–30(–80) cm. Perennials [annuals, subshrubs], 5–150 cm (usually rhizomatous; usually aromatic).
Stems

(sometimes purple-tinged) 1–2+, ± decumbent to ascending or erect, branched.

1 or 2–5+, erect or prostrate to ascending, branched proximally and/or distally, glabrous or hairy (hairs basifixed and/or medifixed, sometimes stellate).

Leaves

basal (soon withering) and cauline; petiolate (bases often clasping) or sessile;

blades ± ovate or elliptic to obovate or spatulate, mostly 7–25+ × 3–5(–10+) cm, 2–3-pinnately lobed (primary lobes mostly 6–24+ pairs, narrowly oblong to linear-elliptic or linear, lobules oblong or ovate to ± lanceolate, sometimes curled), ultimate margins entire or ± dentate, faces usually ± villous or arachno-villous to lanate, sometimes glabrescent or glabrate, usually gland-dotted (in pits).

basal and/or cauline; alternate; petiolate or sessile;

blades mostly obovate to spatulate, usually 1–3-pinnately lobed, ultimate margins entire, crenate, or dentate, faces glabrous or hairy.

Involucres

8–22+ mm diam.

mostly hemispheric or broader, (3–)5–22+ mm diam.

Receptacles

flat to hemispheric.

flat to conic or hemispheric (sometimes hairy), epaleate.

Ray florets

8–21+ (pistillate, fertile; corollas pale yellow to yellow, laminae mostly 1–7+ mm, usually 3-lobed) or 0 (heads quasi-radiant or -radiate or ± disciform, peripheral pistillate florets 15–30+; corollas pale yellow, ± zygomorphic, lobes 3–5, abaxial more pronounced).

usually 10–21+ (pistillate and fertile or neuter; corollas pale yellow to yellow or white, usually with yellowish bases [pink], laminae oblong to flabellate), sometimes 0 (in disciform or quasi-radiate or -radiant heads, peripheral pistillate florets 8–30+; corollas pale yellow, ± zygomorphic, lobes 3–4, sometimes ± raylike).

Disc florets/Disc corollas

(2–)3(–4) mm.

60–300+, bisexual, fertile;

corollas yellow, tubes ± cylindric, throats narrowly funnelform to campanulate, lobes (4–)5, ± deltate.

Phyllaries

persistent, (20–)30–60+ in (2–)3–5+ series, distinct, ± ovate to oblong or oblong to lanceolate or lance-linear (sometimes carinate), unequal, margins and apices (pale to dark brown or blackish) scarious (tips sometimes dilated).

Heads

(2–)5–12(–20+) in corymbiform arrays or borne singly.

usually radiate, sometimes disciform (or quasi-radiate or -radiant), usually in lax to dense, corymbiform arrays, rarely borne singly.

Cypselae

2–3(–4) mm, weakly 5-ribbed or -angled, gland-dotted;

pappi coroniform, 0.1–0.5+ mm (entire or erose to lacerate).

obconic or ± columnar (circular in cross section), ribs (4–)5–10(–12+), faces usually gland-dotted, sometimes glabrous (pericarps without myxogenic cells or resin sacs, embryo sac development tetrasporic);

pappi usually coroniform, rarely 0 [distinct scales or each pappus an adaxial auricle].

x

= 9 (polyploidy).

2n

= 54.

Tanacetum bipinnatum

Tanacetum

Phenology Flowering May–Sep.
Habitat Dunes, other sandy sites, calcareous soils, coastal scrub
Elevation 0–200+ m (0–700+ ft)
Distribution
from FNA
AK; CA; ME; MI; OR; WA; WI; AB; BC; MB; NB; NF; ON; QC; SK; YT; Eurasia
[WildflowerSearch map]
[BONAP county map]
from USDA
North America; Europe; Asia; n Africa; some species widely cultivated
[BONAP county map]
Discussion

The circumscription of Tanacetum bipinnatum adopted here includes not only T. huronense (see E. Hultén 1941–1950, vol. 10, 1968) but T. camphoratum and T. douglasii as well (see D. W. Kyhos and P. H. Raven 1982; C. J. Mickelson and H. H. Iltis 1966). Subspecies bipinnatum has been distinguished from subsp. huronense by having heads borne singly or 2–4 together versus (1–)3–12(–20+) in corymbiform arrays, phyllary margins dark brown versus pale brown, and laminae of ray corollas mostly 3–7 mm versus 1–3 mm. Relatively low plants, 10–20(–40 cm) from dune habitats along the southern shore of Lake Athabasca, Saskatchewan, with mostly 1–4, lanate cauline leaves and 1(–2) heads per flowering stem have been called T. huronense var. floccosum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 160 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves usually not pinnately lobed (sometimes with 1–4+ lateral lobes near bases of blades), ultimate margins ± crenate
T. balsamita
1. Leaves usually 1–3-pinnately lobed, ultimate margins entire or dentate
→ 2
2. Leaf blades 1–2-pinnately lobed (primary lobes 3–5 pairs, ± ovate), faces (at least abaxial) usually puberulent; ray florets 10–21+ (more in "doubles"), corollas white, laminae 2–8(–12+) mm; pappi 0 or coroniform (0.1–0.2+ mm)
T. parthenium
2. Leaf blades 2–3-pinnately lobed (primary lobes 4–24+ pairs, ± oblong to elliptic or linear), faces usually arachno-villous to villous, sometimes glabrescent or glabrous; ray florets 0 or 8–30+, corollas pale yellow to yellow, laminae 1–8+ mm; pappi coroniform (0.1–0.5+ mm)
→ 3
3. Leaves: faces glabrous or sparsely hairy; heads (disciform) 20–200 in corymbiform arrays; involucres 5–10 mm diam
T. vulgare
3. Leaves: faces usually ± villous or arachno-villous to lanate, sometimes glabrescent or glabrate; heads (radiate, quasi-radiant or -radiate, or disciform) (2–)5–12(–20+) in corymbiform arrays or borne singly; involucres 8–22+ mm diam
T. bipinnatum
Source FNA vol. 19, p. 490. FNA vol. 19, p. 489. Author: Linda E. Watson.
Parent taxa Asteraceae > tribe Anthemideae > Tanacetum Asteraceae > tribe Anthemideae
Sibling taxa
T. balsamita, T. parthenium, T. vulgare
Subordinate taxa
T. balsamita, T. bipinnatum, T. parthenium, T. vulgare
Synonyms Chrysanthemum bipinnatum, Chrysanthemum bipinnatum subsp. huronense, T. bipinnatum subsp. huronense, T. camphoratum, T. douglasii, T. huronense, T. huronense var. bifarium, T. huronense var. floccosum, T. huronense var. johannense, T. huronense var. terrae-novae
Name authority (Linnaeus) Schultz-Bipontinus: Tanaceteen, 48. (1844) Linnaeus: Sp. Pl. 2: 843. (1753): Gen. Pl. ed. 5, 366. (1754)
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