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beach sand-spurrey, beach sand-spurry, sticky sand spurry, sticky sand-spurrey

alkali sand-spurrey, alkali sandspurry

Habit Plants strongly perennial with branched, woody base, stout, 5–40 cm, densely stipitate-glandular in inflorescence or throughout. Plants annual, delicate, 5–15 cm, stipitate-glandular throughout or nearly so.
Taproots

becoming stout, woody.

slender.

Stems

erect or ascending to prostrate, usually branched proximally;

main stem 0.8–3 mm diam. proximally.

erect to diffusely spreading, much-branched proximally and distally;

main stem 0.3–0.7 mm diam. proximally.

Leaves

stipules conspicuous, dull white to tan, narrowly triangular, 4.5–11 mm, apex long-acuminate;

blade linear, (0.6–)1–5.5 cm, fleshy, apex apiculate to spine-tipped;

axillary leaves 1–2+ per cluster.

stipules inconspicuous, silvery to dull tan, broadly triangular, 1.5–2 mm, apex acuminate;

blade linear, 0.6–2.3 cm, somewhat fleshy, apex blunt to apiculate;

axillary leaf cluster usually absent.

Pedicels

erect, divergent, or reflexed in fruit.

erect to reflexed in fruit.

Flowers

sepals connate 0.5–1.8 mm proximally, lobes often 3-veined, ovate to lanceolate, 4.5–7 mm, to 8 mm in fruit, margins 0.3–0.7 mm wide, apex blunt to rounded;

petals white or pink to rosy, elliptic to obovate, 0.9–1.3 times as long as sepals in flower;

stamens 9–10;

styles 0.6–3 mm.

sepals connate 0.2–0.5 mm proximally, lobes 1-veined or not, lanceolate to ovate, 2.3–3.1 mm, enlarging little in fruit, margins 0.1–0.3 mm wide, apex obtuse to rounded;

petals white, elliptic to ovate, 0.7–0.8 times sepals;

stamens 4–7;

styles 0.4–0.6 mm.

Capsules

tan, 4.6–10 mm, 0.8–1.4 times as long as sepals.

greenish tan, 2.5–2.8 mm, 0.9–1.2 times sepals.

Seeds

± red-brown, often with submarginal groove or depression, suborbiculate to pyriform, compressed, (0.6–)0.7–1.2 mm, smooth to faintly or prominently tuberculate or sculpturing of parallel, wavy lines or of low, rounded mounds, not papillate;

wing absent or white to reddish brown proximally, 0.1–0.3 mm wide, margins irregular.

black, often with silvery, not iridescent tinge, with submarginal groove, pyriform, somewhat compressed, 0.4–0.6 mm, shiny, sculpturing of low, elongate tubercles, not papillate (40x);

wing absent.

Cymes

simple to 3-compound or flowers solitary and axillary.

simple but commonly 4–8+-compound.

2n

= 18 (Europe).

Spergularia macrotheca

Spergularia diandra

Phenology Flowering spring–fall.
Habitat Sandy beaches, river shores
Elevation 600-700 m (2000-2300 ft)
Distribution
from FNA
CA; OR; WA; BC; including nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; MA; OR; WA; AB; BC; SK; Europe (Mediterranean region); sw Asia; c Asia; n Africa [Introduced in North America; introduced in Australia]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

R. P. Rossbach’s (1940) report of Spergularia diandra from Georgia is referred to 9. S. echinosperma.

The name Spergularia diandra was effectively and validly published via an autographic label distributed in 1851, predating the other commonly seen attributions of this combination.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals pink to rosy; styles 0.6-1.2 mm
var. macrotheca
1. Petals white; styles 1.2-3 mm
→ 2
2. Capsules 1.2-1.4 times as long as sepals; styles 1.2-1.9 mm
var. leucantha
2. Capsules 0.8-1 times as long as sepals; styles 2-3 mm
var. longistyla
Source FNA vol. 5, p. 18. FNA vol. 5, p. 19.
Parent taxa Caryophyllaceae > subfam. Polycarpoideae > Spergularia Caryophyllaceae > subfam. Polycarpoideae > Spergularia
Sibling taxa
S. atrosperma, S. bocconi, S. canadensis, S. diandra, S. echinosperma, S. media, S. platensis, S. rubra, S. salina, S. villosa
S. atrosperma, S. bocconi, S. canadensis, S. echinosperma, S. macrotheca, S. media, S. platensis, S. rubra, S. salina, S. villosa
Subordinate taxa
S. macrotheca var. leucantha, S. macrotheca var. longistyla, S. macrotheca var. macrotheca
Synonyms Arenaria macrotheca Arenaria diandra, S. salsuginea
Name authority (Hornemann ex Chamisso & Schlechtendal) Heynhold: Alph. Aufz. Gew. 689. (1846) (Gussone) Heldreich: Pl. Atticae unnumbered. (1851)
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