Spergularia macrotheca |
Spergularia diandra |
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beach sand-spurrey, beach sand-spurry, sticky sand spurry, sticky sand-spurrey |
alkali sand-spurrey, alkali sandspurry |
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Habit | Plants strongly perennial with branched, woody base, stout, 5–40 cm, densely stipitate-glandular in inflorescence or throughout. | Plants annual, delicate, 5–15 cm, stipitate-glandular throughout or nearly so. | ||||||||
Taproots | becoming stout, woody. |
slender. |
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Stems | erect or ascending to prostrate, usually branched proximally; main stem 0.8–3 mm diam. proximally. |
erect to diffusely spreading, much-branched proximally and distally; main stem 0.3–0.7 mm diam. proximally. |
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Leaves | stipules conspicuous, dull white to tan, narrowly triangular, 4.5–11 mm, apex long-acuminate; blade linear, (0.6–)1–5.5 cm, fleshy, apex apiculate to spine-tipped; axillary leaves 1–2+ per cluster. |
stipules inconspicuous, silvery to dull tan, broadly triangular, 1.5–2 mm, apex acuminate; blade linear, 0.6–2.3 cm, somewhat fleshy, apex blunt to apiculate; axillary leaf cluster usually absent. |
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Pedicels | erect, divergent, or reflexed in fruit. |
erect to reflexed in fruit. |
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Flowers | sepals connate 0.5–1.8 mm proximally, lobes often 3-veined, ovate to lanceolate, 4.5–7 mm, to 8 mm in fruit, margins 0.3–0.7 mm wide, apex blunt to rounded; petals white or pink to rosy, elliptic to obovate, 0.9–1.3 times as long as sepals in flower; stamens 9–10; styles 0.6–3 mm. |
sepals connate 0.2–0.5 mm proximally, lobes 1-veined or not, lanceolate to ovate, 2.3–3.1 mm, enlarging little in fruit, margins 0.1–0.3 mm wide, apex obtuse to rounded; petals white, elliptic to ovate, 0.7–0.8 times sepals; stamens 4–7; styles 0.4–0.6 mm. |
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Capsules | tan, 4.6–10 mm, 0.8–1.4 times as long as sepals. |
greenish tan, 2.5–2.8 mm, 0.9–1.2 times sepals. |
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Seeds | ± red-brown, often with submarginal groove or depression, suborbiculate to pyriform, compressed, (0.6–)0.7–1.2 mm, smooth to faintly or prominently tuberculate or sculpturing of parallel, wavy lines or of low, rounded mounds, not papillate; wing absent or white to reddish brown proximally, 0.1–0.3 mm wide, margins irregular. |
black, often with silvery, not iridescent tinge, with submarginal groove, pyriform, somewhat compressed, 0.4–0.6 mm, shiny, sculpturing of low, elongate tubercles, not papillate (40x); wing absent. |
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Cymes | simple to 3-compound or flowers solitary and axillary. |
simple but commonly 4–8+-compound. |
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2n | = 18 (Europe). |
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Spergularia macrotheca |
Spergularia diandra |
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Phenology | Flowering spring–fall. | |||||||||
Habitat | Sandy beaches, river shores | |||||||||
Elevation | 600-700 m (2000-2300 ft) | |||||||||
Distribution |
CA; OR; WA; BC; including nw Mexico
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ID; MA; OR; WA; AB; BC; SK; Europe (Mediterranean region); sw Asia; c Asia; n Africa [Introduced in North America; introduced in Australia] |
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Discussion | Varieties 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
R. P. Rossbach’s (1940) report of Spergularia diandra from Georgia is referred to 9. S. echinosperma. The name Spergularia diandra was effectively and validly published via an autographic label distributed in 1851, predating the other commonly seen attributions of this combination. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 18. | FNA vol. 5, p. 19. | ||||||||
Parent taxa | Caryophyllaceae > subfam. Polycarpoideae > Spergularia | Caryophyllaceae > subfam. Polycarpoideae > Spergularia | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Arenaria macrotheca | Arenaria diandra, S. salsuginea | ||||||||
Name authority | (Hornemann ex Chamisso & Schlechtendal) Heynhold: Alph. Aufz. Gew. 689. (1846) | (Gussone) Heldreich: Pl. Atticae unnumbered. (1851) | ||||||||
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