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starry campion, widow's frill

bladder campion, common campion, maiden's-tears, silène enflé

Habit Plants perennial; taproot thick; caudex branched. Plants short-lived perennial, glabrous, rarely pubescent, glaucous; taproot stout; caudex woody.
Stems

several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.

several–many, erect, branched and decumbent at base, rarely simple, 20–80 cm.

Leaves

withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;

blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.

mainly cauline, 2 per node, sessile, almost clasping, reduced proximal to inflorescence, blade broadly oblong to oblanceolate or lanceolate, rarely ± linear, 2–8 cm × 5–30 mm, base round, apex acute to acuminate.

Inflorescences

paniculate, open, bracteate, bracteolate, branches elongate, puberulent;

bracts and bracteoles linear-lanceolate, 2–15 mm.

open dichasial cyme, 5–40-flowered, bracteate;

bracts much-reduced, lanceolate.

Pedicels

straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.

0.5–3 cm.

Flowers

calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;

corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;

stamens equaling petals;

styles 3, longer than petals.

bisexual and unisexual, some plants having bisexual flowers, others having pistillate unisexual flowers, 15–20 mm diam.;

calyx pale green, rarely purplish, campanulate, not contracted at mouth or base, inflated, 9–12 mm in flower, 12–18 × 7–11 mm in fruit, herbaceous, papery, venation obscure, reticulate, without conspicuous pale commissures, margins dentate, lobes broadly triangular, 2–3 mm, glabrous;

petals white, ca. 2 times as long as calyx;

limb obovate, emarginate to 2-lobed;

stamens exserted by 2–4 mm;

styles 3, cream to greenish, at most slightly pink tinged, 2 times longer than calyx.

Capsules

globose, opening by 3 broadly triangular teeth;

carpophore 2–3 mm.

ovoid to globose, equaling calyx, opening by 6 teeth;

carpophore 2–3 mm.

Seeds

dark brown, reniform, ca. 1 mm, papillate.

black or nearly so, globose-reniform, 1–1.5 mm, finely tuberculate.

2n

= (34), 48.

= 24.

Silene stellata

Silene vulgaris

Phenology Flowering summer. Flowering summer–fall.
Habitat Rich deciduous woods, river flats, tall-grass prairies Roadsides, waste ground, gravel pits and shores, arable land
Elevation 0-1300 m (0-4300 ft) 0-2000 m (0-6600 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SC; SD; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; YT; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.

Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Silene vulgaris is less variable in North America than in its native Europe, where five subspecies are recognized on the basis of capsule size, petal color, leaf shape, and habit. All North American material appears to belong to subsp. vulgaris, although a few collections from sandy habitats tend to have unusually narrow leaves. Similar plants from Europe have been named var. litoralis (Ruprecht) Jalas and subsp. angustifolia Hayek.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 207. FNA vol. 5, p. 213.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. williamsii, S. wrightii
Synonyms Cucubalus stellatus, S. scabrella, S. stellata var. scabrella Behen vulgaris, S. cucubalus, S. inflata, S. latifolia var. pubescens
Name authority (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) (Moench) Garcke: Fl. N. Mitt.-Deutschland ed. 9, 46. (1869)
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