Poa macrantha |
Poa howellii |
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dune bluegrass, large flower sand dune blue grass, sand-dune blue grass, seashore blue grass |
Howell's blue grass |
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Habit | Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 4 m, stout, robust. | Plants usually annual, rarely longer-lived; densely tufted, tuft bases narrow; not stoloniferous, not rhizomatous. |
Culms | (7)15-60 cm tall, 1.5-2 mm thick, bases decumbent, terete or weakly compressed, smooth or moderately scabrous below the panicles; nodes terete, 0(1) exserted. |
(10) 25-80(120) cm tall, 0.4-1.75 mm thick, usually erect; nodes terete, usually 1-2 exserted. |
Sheaths | closed for about 1/2 their length, terete, glabrous or sparsely retrorsely scabrous, bases of basal sheaths glabrous, distal sheath lengths 1.7-4(6) times blade lengths; collars smooth, glabrous; ligules 1-5 mm, scabrous, truncate to acute, ciliolate; innovation blades to 30 cm, moderately to densely scabrous or hispidulous on and between the veins; cauline blades subequal in length, 2-4 mm wide, involute, thick, somewhat arcuate, firm, abaxial surfaces smooth or moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-10 cm. |
closed for 1/2 - 7/8 their length, usually weakly compressed and keeled, usually scabrous, rarely smooth; ligules 1.5-5(10) mm, smooth or scabrous, acute; blades 1-7(10) mm wide, flat, lax, soft, finely scabrous, apices narrowly prow-shaped, cauline blades 2-10 cm. |
Basal branching | mostly intravaginal, some extravaginal. |
intravaginal. |
Panicles | 3-15 cm, erect, ovoid to lanceolate, contracted, often interrupted, congested, with 15-80 spikelets, rachises glabrous, smooth to moderately scabrous; nodes with 1-2 branches; branches 1-6 cm, erect, stiff, terete to weakly angled, smooth or sparsely to moderately scabrous, with 3-17 spikelets. |
10-25(30) cm, erect, eventually open, with (1)3-5(7) branches per node; branches eventually spreading or reflexed, fairly straight, angled, angles usually moderately to densely scabrous, rarely sparsely scabrous. |
Spikelets | 9-17 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic; florets 3-6(10); rachilla internodes smooth, usually hairy, hairs 0.3-0.4+ mm, rarely glabrous. |
(2)4-6 mm, laterally compressed, with 2-5 florets; rachilla internodes about 1 mm, smooth, usually softly puberulent, infrequently glabrous. |
Glumes | broadly lanceolate, subequal to the adjacent lemmas, distinctly keeled, keels sparsely scabrous near the apices; lower glumes 3-veined; upper glumes usually 7+ mm, 3-5-veined; calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed; lemmas (6)7.5-11 mm, lanceolate, 5-7(11)-veined, distinctly keeled, keels and marginal veins, and sometimes the lateral veins, short-villous to softly puberulent, intercostal regions smooth or scabrous, glabrous or softly puberulent, margins glabrous, apices acute; palea keels scabrous, intercostal regions glabrous; anthers vestigial (0.1-0.2 mm) or (2)3-4(5) mm. |
slightly unequal, lanceolate, distinctly keeled, keels and sometimes the lateral veins sparsely to moderately scabrous; lower glumes 1-3-veined; upper glumes shorter than or subequal to the lowest lemmas; calluses of some or all florets sparsely webbed; lemmas 2.5-3.5 mm, lanceolate to narrowly lanceolate, distinctly keeled, crisply puberulent proximally, hairs evenly distributed, finely scabrous distally, lateral veins obscure to prominent, margins narrowly hyaline, glabrous, apices narrowly acute, infrequently anthocyanic; palea keels sparsely scabrous, glabrous or softly puberulent at midlength, intercostal regions usually softly puberulent; anthers 3, 0.2-1 mm. |
2n | = 28. |
= unknown. |
Poa macrantha |
Poa howellii |
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Distribution |
AK; CA; OR; WA; BC
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CA; OR; WA; BC
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Discussion | Poa macrantha is a dioecious coastal sand dune species that grows from southern Alaska to northern California. It competes better than P. douglasii (see previous) with the invasion of its habitat by Ammophila and other exotic species. It used to be treated as a subspecies of P. douglasii; a few intermediates with that species have been found around the mouth of Little River, California. Although clearly related, the two species are reasonably divergent in a number of characters. Poa macrantha is readily distinguished from P. douglasii by its glabrous rachises and usually longer glumes and lemmas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Poa howellii grows primarily on rocky banks and wooded slopes, from the coastal ranges of southern British Columbia to southern California. It differs from P. bolanderi (see above) in having puberulent, rather than smooth or scabrous, lemmas, and in growing at lower elevations, mostly from near sea level to 1000 m. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 24, p. 551. | FNA vol. 24, p. 534. |
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa |
Sibling taxa | ||
Synonyms | P. douglasii subsp. macrantha | P. bolanderi var. howellii |
Name authority | Vasey | Vasey & Scribn. |
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