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plains bluegrass, prairie speargrass

Bolander's blue grass

Habit Plants perennial; glaucous or not; densely to loosely tufted or the culms solitary, rhizomatous. Plants usually annual, rarely longer-lived; often glaucous; densely tufted, tuft bases narrow, sterile shoots few, not stoloniferous, not rhizomatous.
Culms

15-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-1 exserted.

20-60(70) cm, erect or geniculate at the base;

nodes terete, usually 1-3 exserted.

Sheaths

closed for 1/10 – 1/5(1/4) their length, terete, smooth or sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths (1.2)1.5-9(20) times blade lengths;

ligules (1)1.5-4(5) mm, smooth or sparsely to moderately scabrous, apices obtuse to acute;

blades strongly to gradually reduced in length distally, 1.5-5 mm wide, flat and moderately thin to folded and moderately thick and firm, abaxial surfaces smooth, adaxial surfaces smooth or sparsely to moderately scabrous, primarily over the veins, apices narrowly prow-shaped, flag leaf blades (0.4)1-7(10) cm.

closed for 1/2-1/4 their length, usually compressed and keeled, usually smooth, infrequently scabrous;

ligules 2.5-7 mm, smooth or scabrous, usually decurrent, obtuse to acute;

blades 1.5-5 mm wide, usually flat, rarely folded, lax, soft, smooth or sparsely scabrous, margins scabrous, apices broadly prow-shaped, cauline blades 3-15 cm, flag leaf blades 1-4 cm.

Basal branching

intra- and extravaginal.

both intra- and extravaginal.

Panicles

(2.5)4-12(18) cm, erect, usually narrowly lanceoloid, contracted, sometimes interrupted, infrequently loosely contracted, usually congested, with 25-100 spikelets;

nodes with 1-5 branches;

branches 1-9 cm, erect to infrequently ascending, rarely spreading, terete to weakly angled, smooth or the angles sparsely to moderately scabrous, with 3-24 spikelets.

(5)10-15(25) cm long, 1/4-1/2 the plant height, usually erect, infrequently slightly nodding, usually eventually open, sometimes interrupted, sparse, with 1-3(5) branches per node;

branches initially erect and straight, usually some eventually spreading or reflexed, smooth or sparsely to moderately scabrous.

Spikelets

3.2-7 mm, lengths to 3.5(3.8) times widths, laterally compressed;

florets 2-7;

rachilla internodes smooth, sometimes sparsely puberulent.

(3)4-7 mm, laterally compressed;

florets 2-3(4);

rachilla internodes usually 1-1.2+ mm, smooth or sparsely scabrous, glabrous.

Glumes

lanceolate, distinctly keeled, smooth or sparsely scabrous;

lower glumes 3-veined;

calluses usually glabrous, infrequently webbed, hairs to 1/4 the lemma length;

lemmas 2.5-4.5 mm, lanceolate to narrowly lanceolate, distinctly to weakly keeled, keels and marginal veins short- to long-villous, lateral veins moderately prominent, glabrous or puberulent, intercostal regions usually glabrous, infrequently hairy, hairs to 0.3 mm, margins glabrous, apices acute or blunt;

palea keels scabrous, glabrous or short-villous at midlength, intercostal regions usually glabrous, sometimes puberulent to short-villous;

anthers 1.3-2.2 mm.

unequal, distinctly shorter than the adjacent lemmas, distinctly keeled, keels smooth or sparsely scabrous;

lower glumes 1-3-veined, 1/2 - 2/3 the length of the upper glume, 1/2 - 2/3 the length of the lowest lemmas;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-4 mm, lanceolate to narrowly lanceolate, distinctly keeled, smooth or scabrous throughout, glabrous, lateral veins obscure to moderately prominent, apices narrowly acute, usually anthocyanic near the tip;

palea keels sparsely scabrous;

anthers 3, 0.5-1(1.8) mm.

2n

= 56, 56+1, 56-58, 63, 64, 70, 76, 84, ca. 90, 95+-5, 100, 103.

= 28.

Poa arida

Poa bolanderi

Distribution
from FNA
CO; IA; IL; IN; KS; MI; MN; MT; ND; NE; NJ; NM; OH; OK; SD; TX; WY; AB; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; OR; UT; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa arida grows mainly on the eastern slope of the Rocky Mountains and in the northern Great Plains, primarily in riparian habitats of varying salinity or alkalinity. It is spreading eastward along heavily salted highway corridors. Reports of its occurrence west of the Continental Divide and in southwestern Texas are mostly attributable to misidentifications of P. arctica subsp. aperta (p. 530). P. arctica subsp. grayana (p. 532), and rhizomatous specimens of P. fendleriana (p. 556).

Poa arida may reflect past hybridization between P. secunda (p. 586) and a species of Poa sect. Poa. Poa glaucifolia Scribn. & T.A. Williams refers to specimens of the northern Great Plains that have a more lax growth form with broader leaves and occasionally somewhat open panicles, florets with a small web, and sometimes lacking hairs between the keel and marginal veins of the lemma. Plants with these characteristics have chromosome counts of 2ra = 56 and 70, whereas P. arida sensu stricto usually has 2n = 63, 64, or greater than 70. It is suspected that some of the variability reflects introgression from P. secunda.

Poa arida is a named intersectional hybrid

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa bolanderi grows mainly in pine to fir forest openings of mountain slopes in the western United States, from Washington to California and Utah. It differs from P. howellii (see below) in having smooth to scabrous, rather than puberulent, lemmas; it also grows at higher elevations, mostly at 1500-3000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 599. FNA vol. 24, p. 533.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. glaucifolia P. howellii var. chandleri
Name authority Vasey Vasey
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