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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine

Washoe pine

Habit Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. Trees to 60m; trunk to 1m diam., straight; crown pyramidal.
Bark

red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges.

yellow-brown to reddish, fissured, plates scaly.

Branches

contorted, pendent;

twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

spreading-ascending;

twigs stout, orangish, aging gray, rough.

Buds

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

ovoid, red-brown, 1.5–2cm, not resinous;

scale margins fringed.

Leaves

mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate;

sheath ca. 1cm, soon forming rosette, shed early.

(2–)3 per fascicle, spreading-ascending, persisting (2–)4–6(–7) years, 10–15cm × ca. 1.5mm, slightly twisted, gray-green, all surfaces with stomatal lines, margins finely serrulate, apex acuminate;

sheath 1–2cm, base persistent.

Pollen cones

cylindro-ellipsoid, 7–10mm, purple-red.

cylindric, 10–20mm, red-purple.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile;

apophyses much thickened, sharply keeled;

umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale.

cones maturing in 2 years, shedding seeds soon thereafter, not persistent, spreading, slightly asymmetric, ovoid-conic before opening, broadly ovoid when open, 7–10cm, tan or pale red-brown, sessile, abaxial surface of scales darker and sharply contrasting in color with adaxial surface;

apophyses slightly raised, low pyramidal;

umbo central, narrowly pyramidal, tapering into short, reflexed, fine prickle.

2n

=24.

Pinus longaeva

Pinus washoensis

Habitat Subalpine and alpine Dry montane forests
Elevation 1700–3400m (5600–11200ft) 2100–2500m (6900–8200ft)
Distribution
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
Discussion

Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus washoensis often occurs in large stands and resembles P. jeffreyi. The number and posture of seed-cone scales fall within the ranges given for P. jeffreyi. The abaxial surface of these scales has a significantly darker pigmentation, however; such a color contrast is not apparent in P. jeffreyi. Forest geneticists have developed hybrids between P. washoensis and related yellow pines, but no natural hybrids have been observed. Some workers regard P. washoensis as closely related to—or even conspecific with--- P. ponderosa.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana
Synonyms P. aristata var. longaeva
Name authority D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) H. Mason & Stockwell: Madroño 8: 62. (1945)
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