Paysonia lasiocarpa
Paysonia
roughpod bladderpod
decumbent to procumbent, (slender), 1–5 dm.
erect, ascending, decumbent, or procumbent, unbranched or branched.
basal and cauline;
petiolate or sessile;
basal rosulate, petiolate, blade margins entire, dentate, or pinnately lobed;
cauline sessile, blade (base usually auriculate or sagittate, sometimes amplexicaul), margins dentate to lobed.
blade (oblanceolate), 3–10 cm × 10–30 mm, margins sinuate-dentate to somewhat lobed or incised, surfaces densely pubescent.
blade obovate-elliptic to oblong, 1–4 cm × 5–20 mm, base cuneate to auriculate (narrowed toward base), margins sinuate-dentate to incised, (surfaces densely pubescent).
(several-flowered), often strongly elongated in fruit, (rachis straight).
sepals (green), 4–5.5 × 1.5–2 mm, (nearly acuminate), pubescent or hirsute (trichomes branched);
petals light yellow (often drying purplish), 5–9 × 4–6 mm, (claw relatively short), apex rounded;
filaments not dilated basally, (2.5–3.5 mm, anthers sagittate, 2–3 mm, glands with horn-like projections at base of filaments).
sepals (deciduous), suberect, ascending, or spreading, oblong, (equal), lateral pair not saccate basally, (margins membranous);
petals yellow or white, broadly obovate, claw slightly differentiated from blade, (apex rounded, truncate, retuse, or emarginate);
stamens (erect), tetradynamous;
filaments dilated or not basally;
anthers oblong or sagittate, (not apiculate);
nectar glands confluent, subtending bases of median stamens, surrounding lateral stamens.
recurved, slightly curved, 1–3 cm, (racemes lax in fruit), densely pubescent.
ascending to divaricate or recurved, (straight or slightly curved), slender.
sessile, orbicular, elliptic, or cordate, 5–9 × 4–9 mm, (angustiseptate or slightly so);
valves (rounded or strongly keeled), densely pubescent, trichomes branched, sometimes with simple, larger ones;
replum oblong to nearly elliptical;
septum complete, (transparent);
ovules 14–32 per ovary, (attached in proximal 1/2);
style 1–1.5 mm;
stigma expanded.
silicles, subsessile or sessile, globose, subglobose, subpyriform, suborbicular, orbicular, elliptic, cordate, or obovoid, terete, flattened, latiseptate, or angustiseptate;
valves (papery or leathery), not veined, glabrous or pubescent, (trichomes branched and/or simple);
replum not flattened;
septum complete, perforated, or reduced to a rim, (membranous);
ovules 4–40 per ovary;
style distinct, (persistent, slender);
stigma capitate, entire.
suborbicular (slightly longer than broad), 1–2.2 mm.
sub-biseriate, flattened, margined, orbicular, suborbicular, or oval;
cotyledons accumbent.
= 7, 8, 9.
Paysonia lasiocarpa
Paysonia
Subspecies 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species 8 (8 in the flora).
Paysonia was segregated from Lesquerella (now Physaria) based on its different base chromosome number, presence of auriculate stem leaves, typically annual duration, trichomes that are neither stellate nor tuberculate, and phylogenetic analysis of DNA sequences. Some species of the genus have very limited distributions and are of conservation concern. Five (P. densipila, P. lescurii, P. lyrata, P. perforata, and P. stonensis) are known to be interfertile under experimental conditions, and some combinations form fertile hybrids in the field (R. C. Rollins 1988).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Annuals; trichomes simple and branched; fruits inflated, valves rounded. | subsp. lasiocarpa |
1. Annuals, biennials, or perennials; trichomes usually simple, sometimes mixed with smaller, branched ones; fruits not inflated, valves somewhat to strongly keeled. | subsp. berlandieri |
1. Fruits orbicular, elliptic, or cordate | → 2 |
1. Fruits globose, subglobose, subpyriform, or obovoid | → 3 |
2. Fruits angustiseptate; fruiting pedicels recurved; cauline leaf blades: base cuneate to auriculate, not amplexicaul; filaments not dilated basally. | P. lasiocarpa |
2. Fruits latiseptate; fruiting pedicels divaricate or nearly straight; cauline leaf blades: base auriculate to sagittate, sometimes amplexicaul; filaments dilated basally. | P. lescurii |
3. Petals white or pale lavender; septums perforate or nearly absent | → 4 |
3. Petals yellow; septums complete | → 5 |
4. Fruits subpyriform, valves sparsely hirsute or glabrate; styles glabrous. | P. perforata |
4. Fruits subglobose, valves densely hirsute; styles hirsute, at least proximally. | P. stonensis |
5. Fruit valves pubescent; styles glabrous or pubescent proximally. | P. densipila |
5. Fruit valves and styles glabrous | → 6 |
6. Fruits 3-4 mm; ovules 4-8 per ovary; septums opaque. | P. lyrata |
6. Fruits 4-6(-8) mm; ovules (8-)12-28(-40) per ovary; septums translucent | → 7 |
7. Stems hirsute proximally, trichomes simple; petals 4-5 mm wide. | P. auriculata |
7. Stems pubescent proximally, trichomes branched; petals 6-9 mm wide. | P. grandiflora |
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