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fall panicgrass, fall panicum, knee grass, panic d'automne, smooth witchgrass

broom-corn millet, broomcorn, hog millet, millet, millet commun, panic millet, proso millet

Habit Plants annual or short-lived perennials in the Flora region, perennial in the tropics; usually terrestrial, sometimes aquatic but not floating. Plants annual; sometimes branching from the lower nodes.
Culms

5-200 cm tall, 0.4-3 mm thick, decumbent to erect, commonly geniculate to ascending, rooting at the lower nodes when in water, simple to divergently branched from the lower and middle nodes, usually succulent, slightly compressed, glabrous;

nodes usually swollen, sometimes constricted on robust plants, glabrous;

internodes glabrous, shiny, pale green to purplish.

20-210 cm, stout, not woody;

nodes puberulent;

internodes usually with papillose-based hairs, sometimes nearly glabrous, not succulent.

Sheaths

compressed, inflated, sparsely pubescent near the base, elsewhere mostly glabrous, sparsely pilose, or hispid, hairs sometimes papillose-based, margins or throat ciliate, with papillose-based hairs;

ligules 0.5-2 mm;

blades 10-65 cm long, 3-25 mm wide, glabrous or sparsely pilose, often scabrous near the margins, midribs stout, whitish.

Leaves

numerous;

sheaths terete, densely pilose, with papillose-based and caducous hairs;

ligules membranous, ciliate, cilia 1-3 mm;

blades 15-40 cm long, 7-25 mm wide.

Panicles

4-40 cm, diffuse, lax, with a few spikelets;

branches to 15 cm, alternate or opposite, occasionally verticillate, ascending to spreading, stiff, scabrous;

pedicels 1-6 mm, sharply 3-angled, scabrous, expanded to cuplike apices, appressed mostly to the abaxial side of the branches.

6-20 cm long, 4-11 cm wide, included or shortly exserted at maturity, dense;

branches stiff, appressed to spreading, spikelets solitary, confined to the distal portions;

pedicels 1-9 mm, scabrous and sparsely pilose.

Spikelets

1.8-3.8 mm long, 0.7-1.2 mm wide, ellipsoid to narrowly ovoid, light green to red-purple, glabrous, acute to acuminate.

4-6 mm, ovoid, usually glabrous.

Lower glumes

0.6-1.2 mm, 1/4 - 1/3 as long as the spikelets, 0-3-veined, obtuse to acute;

upper glumes and lower lemmas similar, exceeding the upper florets by 0.3-0.6 mm, 7-9-veined;

lower paleas vestigial to almost as long as the lower lemmas;

lower florets sterile;

upper florets 1.4-2.5 mm long, 0.7-1.1 mm wide, narrowly ellipsoid, smooth, shiny, stramineous to nigrescent, with pale veins.

2.8-3.6 mm, 1/2 - 3/4 as long as the spikelets, 5-7-veined, veins scabridulous distally, apices attenuate;

upper glumes 4-5.1 mm, slightly exceeding the upper florets, 11-13(15)-veined, veins scabridulous distally;

lower florets sterile;

lower lemmas 4-4.8 mm, slightly exceeding the upper florets, 9-13-veined, veins scabridulous distally;

lower paleas 1.2-1.6 mm, 1/2 or less the length of the upper florets, truncate to bilobed;

upper florets 3-3.8 mm long, 2-2.5 mm wide, smooth or striate, more or less shiny, stramineous to orange, red-brown, or blackish, persisting in the spikelets or disarticulating at maturity.

2n

= 36, 54.

= 36, 40, 42, 49, 54, 72.

Panicum dichotomiflorum

Panicum miliaceum

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; HI; PR; BC; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WY; HI; PR; AB; BC; LB; MB; NB; NS; ON; PE; QC; SK; Virgin Islands
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum dichotomiflorum grows in open, often wet, disturbed areas such as cultivated and fallow fields, roadsides, ditches, open stream banks, receding shores, clearings in flood plain woods, and sometimes in shallow water. It is probably native throughout the eastern United States and adjacent Canada, but introduced elsewhere, including in the western United States. Its size and habit may be partly under genetic control, but these features also seem to be strongly affected by moisture levels, soil richness, competition, and the time of germination.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum miliaceum is native to Asia, where it has been cultivated for thousands of years. In the Flora region, it is grown for bird seed and is occasionally planted for game birds. It is also found in corn fields and along roadsides. In Asia, P. miliaceum is still grown for fodder and as a cereal, its fast germination and short growth period enabling it to be sown following a spring crop. It also has one of the lowest water requirements of any cereal grain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets 1.8-2.2 mm long, widest at the middle, acute; upper glumes and lower lemmas submembranaceous; pedicels often over 3 mm long
subsp. puritanorum
1. Spikelets 2.2-3.8 mm long, widest below the middle, acuminate; upper glumes and lower lemmas subcoriaceous; most pedicels less than 3 mm long.
→ 2
2. Sheaths glabrous or sparsely pilose, hairs not papillose-based
subsp. dichotomiflorum
2. Sheaths hispid, hairs papillose-based
subsp. bartowense
1. Mature upper florets blackish, disarticulating at maturity; culms 70-210 cm tall; panicles erect, exserted at maturity, about twice as long as wide; panicle branches ascending to spreading; pulvini well-developed
subsp. ruderale
1. Mature upper florets stramineous to orange, not disarticulating; culms 20-120 cm tall; panicles usually nodding, not fully exserted, more than twice as long as wide; panicle branches ascending to appressed; pulvini almost absent
subsp. miliaceum
Source FNA vol. 25, p. 469. FNA vol. 25, p. 456.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Dichotomiflora Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Panicum
Sibling taxa
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
Subordinate taxa
P. dichotomiflorum subsp. bartowense, P. dichotomiflorum subsp. dichotomiflorum, P. dichotomiflorum subsp. puritanorum
P. miliaceum subsp. miliaceum, P. miliaceum subsp. ruderale
Name authority Michx. L.
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