Leucospora
Leucospora multifida
leucospora, paleseed
narrow-leaf paleseed, obe-wan-conobea
erect or ascending, glandular-villosulous.
(4–)10–30(–41) cm, usually much-branched from base;
branches spreading to ascending.
cauline, opposite or whorled;
petiole present;
blade not fleshy, not leathery, margins pinnatifid to bipinnatifid.
2 or 3(or 4) per node;
blade ovate to triangular-ovate in outline, rarely lanceolate in outline, 8–30 × 4–23 mm, ultimate segments lanceolate to linear.
axillary, flowers solitary;
bracts absent.
present;
bracteoles absent.
spreading to ascending, (1–)2–5 mm in flower, to 7 mm in fruit, glandular-villosulous.
bisexual;
sepals 5, distinct, lanceolate to linear-lanceolate, calyx obscurely bilaterally symmetric, tubular;
corolla pale blue to pale lavender, bilaterally symmetric, bilabiate, tubular, tube base not spurred or gibbous, throat not densely pilose internally, lobes 5, abaxial 3, adaxial 2;
stamens 4, medially adnate to corolla, didynamous, filaments glabrous;
staminode 0;
ovary 2-locular, placentation axile;
stigma cuneate.
calyx lobes slightly falcate, abaxial slightly shorter than adaxial, 2.6–4 × 0.4–0.7 mm;
corolla glabrous, tube 2–3 mm, throat open, yellow or greenish yellow abaxially within, lobes 1 × 1 mm;
stamens included;
style 0.5–1 mm.
capsules, dehiscence septicidal.
ovoid, 4.5 × 1.8–2.3 mm.
50–100, white, parallel-ridged, ovoid, wings absent.
0.3–0.4 mm, 8-ribbed, ribs longitudinal, straight.
Leucospora
Leucospora multifida
Species 2 (1 in the flora).
The second species of Leucospora, as circumscribed here, is L. coahuilensis Henrickson, which is known from Chihuahua, Coahuila, Durango, and Zacatecas, Mexico. It differs from L. multifida in its longer corollas, shorter pedicels, longer and more slender capsules, and smaller seeds (J. Henrickson 1989).
Based on a molecular phylogenetic study, D. Estes and R. L. Small (2008) suggested that Leucospora belongs to a clade that includes Bacopa, Gratiola, Mecardonia, Scoparia, and Stemodia, genera traditionally placed in Gratioleae. Relationships of some species, and the circumscriptions of some genera and the tribe itself, remain uncertain due to limited sampling. Leucospora was found to be sister to Stemodia verticillata (Miller) Hassler; Stemodia was paraphyletic, with the type species of that genus, S. maritima Linnaeus, included in a second, separate clade. B. L. Turner and C. C. Cowan (1993) included Leucospora in Stemodia based on morphological evidence.
Schistophragma, another genus possibly allied with Leucospora, has not been examined molecularly; it shares some morphological similarities with Leucospora. On the basis of morphology, J. Henrickson (1989) argued that Schistophragma should be submerged in Leucospora. Pending more complete taxon sampling for Gratioleae, Leucospora is retained here in its traditional sense.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
As noted by F. W. Pennell (1935), populations of Leucospora multifida often are associated with calcareous substrates; populations also occur on other substrates. They occasionally are reported along railroads and in damp sites in parking lots in urban areas.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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