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leucospora, paleseed

Habit Herbs, annual.
Stems

erect or ascending, glandular-villosulous.

Leaves

cauline, opposite or whorled;

petiole present;

blade not fleshy, not leathery, margins pinnatifid to bipinnatifid.

Inflorescences

axillary, flowers solitary;

bracts absent.

Pedicels

present;

bracteoles absent.

Flowers

bisexual;

sepals 5, distinct, lanceolate to linear-lanceolate, calyx obscurely bilaterally symmetric, tubular;

corolla pale blue to pale lavender, bilaterally symmetric, bilabiate, tubular, tube base not spurred or gibbous, throat not densely pilose internally, lobes 5, abaxial 3, adaxial 2;

stamens 4, medially adnate to corolla, didynamous, filaments glabrous;

staminode 0;

ovary 2-locular, placentation axile;

stigma cuneate.

Fruits

capsules, dehiscence septicidal.

Seeds

50–100, white, parallel-ridged, ovoid, wings absent.

Leucospora

Distribution
from USDA
e North America; n Mexico; c North America
[BONAP county map]
Discussion

Species 2 (1 in the flora).

The second species of Leucospora, as circumscribed here, is L. coahuilensis Henrickson, which is known from Chihuahua, Coahuila, Durango, and Zacatecas, Mexico. It differs from L. multifida in its longer corollas, shorter pedicels, longer and more slender capsules, and smaller seeds (J. Henrickson 1989).

Based on a molecular phylogenetic study, D. Estes and R. L. Small (2008) suggested that Leucospora belongs to a clade that includes Bacopa, Gratiola, Mecardonia, Scoparia, and Stemodia, genera traditionally placed in Gratioleae. Relationships of some species, and the circumscriptions of some genera and the tribe itself, remain uncertain due to limited sampling. Leucospora was found to be sister to Stemodia verticillata (Miller) Hassler; Stemodia was paraphyletic, with the type species of that genus, S. maritima Linnaeus, included in a second, separate clade. B. L. Turner and C. C. Cowan (1993) included Leucospora in Stemodia based on morphological evidence.

Schistophragma, another genus possibly allied with Leucospora, has not been examined molecularly; it shares some morphological similarities with Leucospora. On the basis of morphology, J. Henrickson (1989) argued that Schistophragma should be submerged in Leucospora. Pending more complete taxon sampling for Gratioleae, Leucospora is retained here in its traditional sense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 270. Author: Craig C. Freeman.
Parent taxa Plantaginaceae
Subordinate taxa
L. multifida
Name authority Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 87. (1834)
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