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downy buckwheat, red creek buckwheat, red creek Howell's-buckwheat

Howell's-buckwheat

Habit Plants weakly erect, grayish to greenish, 0.5–3 × 1–2 dm, mostly densely silky-puberulent. Herbs, annual; taproot slender.
Stems

0.3–0.8 dm.

arising directly from the root, weakly erect to spreading, solid, not fistulose or disarticulating into ringlike segments, silky-puberulent.

Leaves

blades obovate to rounded, 0.5–1.5 × 0.5–1.5 cm.

usually quickly deciduous, basal, rosulate;

petiole present;

blade obovate to rounded or somewhat reniform, margins entire.

Inflorescences

spreading, open, 5–25 cm;

bracts linear to narrowly lanceolate, 2–5(–9) × 1–2(–2.5) mm, somewhat foliaceous, puberulent.

terminal, cymose;

branches dichotomous or trichotomous at proximal node, otherwise dichotomous, not brittle or disarticulating into segments, round, silky-puberulent;

bracts 3, connate proximally, triangular or linear to narrowly lanceolate, scalelike or somewhat leaflike, not awned, glabrous to puberulent.

Peduncles

absent.

Flowers

perianth white to pale yellow, becoming rose or red in fruit, 1.5–2 mm, minutely pustulose basally and along midribs;

tepals slightly dimorphic, those of outer whorl narrowly ovate, slightly auriculate basally with undulate-crisped margins in fruit, those of inner whorl narrowly oblanceolate, often shorter than those of outer whorl;

filaments 0.8–1.2 mm.

4–7 per involucral cluster;

perianth white or pale yellowish to rose or red, broadly campanulate when open, narrowly urceolate when closed, smooth or minutely pustulose abaxially, glabrous;

tepals 6, connate 1/4 their length, monomorphic or slightly dimorphic, entire apically;

stamens 9;

filaments basally adnate, glabrous;

anthers white to pale pink, oblong.

Achenes

1–1.5 mm.

included, light brown, not winged, 3-gonous, glabrous.

Seeds

embryo curved.

Involucral

bracts oblanceolate, 0.5–1.2 × 0.1–0.3 mm, villous abaxially, outermost one 2- or 3-lobed apically.

bracts obscure, in tight spiral of (3–)4(–7) distinct, oblanceolate to obovate lobes, bractlike, not awned.

Johanneshowellia puberula

Johanneshowellia

Phenology Flowering May–Sep.
Habitat Sandy flats and slopes, saltbush and sagebrush communities, pinyon-juniper woodlands
Elevation (500-)800-2800 m ((1600-)2600-9200 ft)
Distribution
from FNA
CA; NV; UT
[BONAP county map]
from USDA
w United States
[BONAP county map]
Discussion

Johanneshowellia puberula is infrequent to occasionally locally common. Populations are widely scattered from the Cottonwood Mountains of Inyo County, California, across northern Clark, southern Eureka, Lincoln, Nye, and White Pine counties of Nevada, into Beaver, Iron, Millard, and Washington counties in Utah.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 2 (2 in the flora).

With recognition of Johanneshowellia, all members of Eriogonum have their involucral bracts fused into a distinct, turbinate to campanulate or hemispheric, tubular structure. In Johanneshowellia, the four to seven, distinct, involucral bracts are arranged in a tight spiral. Each 0.5–1.2 mm, awnless, entire to deeply lobed bract subtends a flower-bearing pedicel. These involucral bracts are themselves usually obscured by inflorescence bracts that subtend each node. As a result, care must be taken to observe the true nature of the involucral complex. The genus is allied with Eriogonum subg. Oregonium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perianths minutely pustulose basally and along midribs, 1.5-2 mm in fruit; outer tepals narrowly ovate, slightly auriculate basally with slightly undulate-crisped margins in fruit; plants grayish to greenish, mostly densely silky-puberulent
J. puberula
1. Perianths smooth, 2-2.5 mm in fruit; tepals lanceolate, not auriculate basally, margins entire; plants reddish, thinly silky-puberulent
J. crateriorum
Source FNA vol. 5, p. 443. FNA vol. 5, p. 443. Author: James L. Reveal.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Johanneshowellia Polygonaceae > subfam. Eriogonoideae
Sibling taxa
J. crateriorum
Subordinate taxa
J. crateriorum, J. puberula
Synonyms Eriogonum puberulum, Eriogonum puberulum var. venosum Eriogonum unranked Puberula
Name authority (S. Watson) Reveal: Brittonia 56: 302. (2004) Reveal: Brittonia 56: 299. (2004)
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