Johanneshowellia |
Johanneshowellia puberula |
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Howell's-buckwheat |
downy buckwheat, red creek buckwheat, red creek Howell's-buckwheat |
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Habit | Herbs, annual; taproot slender. | Plants weakly erect, grayish to greenish, 0.5–3 × 1–2 dm, mostly densely silky-puberulent. | ||||
Stems | arising directly from the root, weakly erect to spreading, solid, not fistulose or disarticulating into ringlike segments, silky-puberulent. |
0.3–0.8 dm. |
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Leaves | usually quickly deciduous, basal, rosulate; petiole present; blade obovate to rounded or somewhat reniform, margins entire. |
blades obovate to rounded, 0.5–1.5 × 0.5–1.5 cm. |
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Inflorescences | terminal, cymose; branches dichotomous or trichotomous at proximal node, otherwise dichotomous, not brittle or disarticulating into segments, round, silky-puberulent; bracts 3, connate proximally, triangular or linear to narrowly lanceolate, scalelike or somewhat leaflike, not awned, glabrous to puberulent. |
spreading, open, 5–25 cm; bracts linear to narrowly lanceolate, 2–5(–9) × 1–2(–2.5) mm, somewhat foliaceous, puberulent. |
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Peduncles | absent. |
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Flowers | 4–7 per involucral cluster; perianth white or pale yellowish to rose or red, broadly campanulate when open, narrowly urceolate when closed, smooth or minutely pustulose abaxially, glabrous; tepals 6, connate 1/4 their length, monomorphic or slightly dimorphic, entire apically; stamens 9; filaments basally adnate, glabrous; anthers white to pale pink, oblong. |
perianth white to pale yellow, becoming rose or red in fruit, 1.5–2 mm, minutely pustulose basally and along midribs; tepals slightly dimorphic, those of outer whorl narrowly ovate, slightly auriculate basally with undulate-crisped margins in fruit, those of inner whorl narrowly oblanceolate, often shorter than those of outer whorl; filaments 0.8–1.2 mm. |
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Achenes | included, light brown, not winged, 3-gonous, glabrous. |
1–1.5 mm. |
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Seeds | embryo curved. |
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Involucral | bracts obscure, in tight spiral of (3–)4(–7) distinct, oblanceolate to obovate lobes, bractlike, not awned. |
bracts oblanceolate, 0.5–1.2 × 0.1–0.3 mm, villous abaxially, outermost one 2- or 3-lobed apically. |
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Johanneshowellia |
Johanneshowellia puberula |
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Phenology | Flowering May–Sep. | |||||
Habitat | Sandy flats and slopes, saltbush and sagebrush communities, pinyon-juniper woodlands | |||||
Elevation | (500-)800-2800 m ((1600-)2600-9200 ft) | |||||
Distribution |
w United States |
CA; NV; UT |
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Discussion | Species 2 (2 in the flora). With recognition of Johanneshowellia, all members of Eriogonum have their involucral bracts fused into a distinct, turbinate to campanulate or hemispheric, tubular structure. In Johanneshowellia, the four to seven, distinct, involucral bracts are arranged in a tight spiral. Each 0.5–1.2 mm, awnless, entire to deeply lobed bract subtends a flower-bearing pedicel. These involucral bracts are themselves usually obscured by inflorescence bracts that subtend each node. As a result, care must be taken to observe the true nature of the involucral complex. The genus is allied with Eriogonum subg. Oregonium. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Johanneshowellia puberula is infrequent to occasionally locally common. Populations are widely scattered from the Cottonwood Mountains of Inyo County, California, across northern Clark, southern Eureka, Lincoln, Nye, and White Pine counties of Nevada, into Beaver, Iron, Millard, and Washington counties in Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 443. | FNA vol. 5, p. 443. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Eriogonum unranked Puberula | Eriogonum puberulum, Eriogonum puberulum var. venosum | ||||
Name authority | Reveal: Brittonia 56: 299. (2004) | (S. Watson) Reveal: Brittonia 56: 302. (2004) | ||||
Web links |