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leatherstem, limberbush, sangre de drago, sangregado

bahada, jicamilla, jirawilla, ragged jatropha, ragged nettlespurge

Habit Subshrubs, to 1 m, dioecious, rhizomatous, often forming colonies. Herbs, perennial, to 0.5 m, monoecious, with ± fleshy subterranean caudex to 11 cm diam., stem and root scars round.
Stems

spreading, reddish brown (when actively growing) or red to dark red or grayish (when dormant), much-branched, rubbery-succulent, glabrous; short shoots common;

latex watery, colorless to cloudy-whitish in fast growing shoots, blood red in basal portion of older shoots and rhizomes.

erect, green, usually sparsely branched, herbaceous, somewhat succulent, glabrous; short shoots absent;

latex watery, colorless.

Leaves

deciduous, fascicled on short shoots;

stipules deciduous, linear, 1–2 mm, undivided;

petiole 0–0.2 cm, not stipitate-glandular;

blade linear-spatulate to narrowly obovate, 1.5–5(–7) × 0.2–0.7 cm, usually unlobed, sometimes shallowly, rarely deeply, 3-lobed, base attenuate, margins entire, apex rounded, membranous, surfaces glabrous;

venation pinnate (palmate if lobed).

± evenly distributed on long shoots;

stipules persistent, 4–10 mm, filiform-divided;

petiole 4.8–10.5(–13.5) cm, not stipitate-glandular;

blade cordate in outline, 11–16 × 9.3–11.2 cm, (3–)5–7(–9)-lobed to middle, base cordate, margins coarsely dentate, apex acuminate, membranous, abaxial surface glabrous, adaxial glabrous, except puberulent on veins, margins sometimes puberulent or with setae;

venation palmate.

Inflorescences

terminal or axillary, fascicles;

peduncle absent;

bracts absent.

bisexual, terminal and subterminal, cymes;

peduncle 1.5–4.5 cm;

bracts (6–)8–16 mm, margins serrate, glabrous.

Pedicels

1–2.5 mm.

2.5–4 mm.

Staminate flowers

sepals distinct or connate basally, ovate to elliptic-lanceolate, 2.5–3 × 1.2–1.4 mm, margins entire, apex acute, abaxial surface hairy, adaxial glabrous;

corolla white to pinkish white, urceolate-tubular, petals connate most of length, 5–6 × 1.4–1.5 mm, abaxial surface sparsely puberulent abaxially, adaxial glabrous;

stamens 10 in 2 whorls (5 + 5);

filaments of both whorls connate from 1/5–3/4+ length, outer whorl 2–3 mm, inner whorl 3–5 mm.

sepals distinct or connate to 1/2 length, lanceolate, 5–7 × 1–2 mm, margins deeply divided, apex acuminate, surfaces glabrous;

corolla light pink, often with white striations, rotate, petals distinct or connate 1/4 length, 8–11.5 × 2.5–4.5 mm, surfaces glabrous;

stamens 8(–9) in 2 whorls (5 + 3);

filaments of outer whorl distinct, of inner whorl connate to 1/2 length, outer whorl 3.5–6 mm, inner whorl 4.5–9 mm.

Pistillate flowers

resembling staminate;

carpel 1;

style 2.5–3 mm.

resembling staminate, but slightly larger;

carpels 3;

styles distinct or connate only at base, 3.5–4 mm.

Capsules

ellipsoidal, 1.2–1.4 × 1.1–1.3 cm, tardily dehiscent.

± spheric, 1.2–1.3 × 1.2–1.3 cm, distinctly 3-lobed, explosively dehiscent.

Seeds

solid gray-brown, spheric, 12 mm diam.;

caruncle rudimentary.

pale gray, ellipsoidal, 8–9 × 6–6.5 mm;

caruncle prominent.

2n

= 44.

= 22.

Jatropha dioica

Jatropha macrorhiza

Phenology Flowering and fruiting spring–summer.
Habitat Hillsides, mesas, sandy washes.
Elevation 1000–2600 m. (3300–8500 ft.)
Distribution
from FNA
TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

R. McVaugh (1945b) recognized two varieties in Jatropha dioica based primarily on leaf shape; the two are not readily recognizable and probably hybridization between them has resulted in intermediate populations, as McVaugh noted. The varieties are treated here with some reservation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Jatropha macrorhiza is known in Texas only from a single collection made in Presidio County in 1938; it is common in nearby Chihuahua and appears to be native to trans-Pecos Texas, but may now be extirpated there.

Plants from the United States often have been called var. septemfida, named in reference to having seven (as opposed to five) leaf lobes or segments. However, leaf segment number varies, even on the same plant, from four to nine (usually five to seven). Hence, a distinct variety cannot be recognized. R. McVaugh (1945) reported this species to have ten stamens; I have not seen any live or herbarium specimens with ten; two collections have recorded nine.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades oblanceolate to narrowly obovate, 3.5–5(–7) cm; sepals distinct; stamen filaments connate 3/4+ length; petals pinkish white.
var. dioica
1. Leaf blades linear-spatulate, 1.5–2.5 cm; sepals connate basally; stamen filaments connate to 1/4 length; petals white.
var. graminea
Source FNA vol. 12, p. 204. FNA vol. 12, p. 201.
Parent taxa Euphorbiaceae > Jatropha Euphorbiaceae > Jatropha
Sibling taxa
J. canescens, J. cardiophylla, J. cathartica, J. cuneata, J. curcas, J. gossypiifolia, J. integerrima, J. macrorhiza, J. multifida
J. canescens, J. cardiophylla, J. cathartica, J. cuneata, J. curcas, J. dioica, J. gossypiifolia, J. integerrima, J. multifida
Subordinate taxa
J. dioica var. dioica, J. dioica var. graminea
Synonyms J. arizonica, J. macrorhiza var. septemfida
Name authority Sessé: Gaz. Lit. México 3(suppl.): 4. (1794) Bentham: Pl. Hartw., 8. (1839)
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