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willow feather-moss

Habit Plants in ± stiff mats, yellowish to dark green, dull.

leaves ovate-lanceolate to ovate-triangular;

basal laminal cells short-rectangular, often pigmented, walls firm, eporose, region weakly differentiated;

medial cells 2–7:1.


dark red, 1.2–2.2 cm.


constricted below mouth when dry;

exothecial cells long- to short-rectangular, 30–105 × 18–35 µm, walls firm, not collenchymatous;

annulus separating, 2–4-seriate, cell walls thin;

operculum acute, apiculate, or rostrate;

exostome teeth yellow-brown, external surface cross striolate basally, coarsely papillose distally, internal surface trabeculate;

endostome basal membrane high, segments not or narrowly perforate.


finely papillose.

Amblystegium varium

from USDA
North America; Mexico; Central America; South America; Eurasia; s Africa; Pacific Islands (New Zealand); Antarctica

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

1. Plants small to moderately large; stems to 10 cm; costae narrower than 100 µm at base, to mid leaf, percurrent, or rarely short-excurrent; lamina 1-stratose.
subsp. varium
1. Plants extremely large; stems to 15 cm; costae 110-140 µm wide at base, excurrent; lamina sometimes partially 2-stratose, especially at base near costa.
subsp. noterophilum
Source FNA vol. 28, p. 303.
Parent taxa Amblystegiaceae > Hygroamblystegium
Subordinate taxa
H. varium subsp. noterophilum, H. varium subsp. varium
Synonyms Leskea varia, Amblystegium varium
Name authority (Hedwig) Mönkemeyer: Hedwigia 50: 275. (1911)
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