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willow feather-moss

Photo is of parent taxon
Habit Plants in ± stiff mats, yellowish to dark green, dull. Plants small to moderately large.
Stem(s)

leaves ovate-lanceolate to ovate-triangular;

basal laminal cells short-rectangular, often pigmented, walls firm, eporose, region weakly differentiated;

medial cells 2–7:1.

leaves 0.6–2 × 0.3–0.8 mm, lamina 1-stratose;

margins entire or nearly so to serrate;

costa to mid leaf, percurrent, or rarely short-excurrent, narrower than 100 µm at base;

medial laminal cells 10–60 × 5–15 µm.

Seta

dark red, 1.2–2.2 cm.

Capsule

constricted below mouth when dry;

exothecial cells long- to short-rectangular, 30–105 × 18–35 µm, walls firm, not collenchymatous;

annulus separating, 2–4-seriate, cell walls thin;

operculum acute, apiculate, or rostrate;

exostome teeth yellow-brown, external surface cross striolate basally, coarsely papillose distally, internal surface trabeculate;

endostome basal membrane high, segments not or narrowly perforate.

Spores

finely papillose.

Amblystegium varium

Hygroamblystegium varium subsp. varium

Distribution
from USDA
North America; Mexico; Central America; South America; Eurasia; s Africa; Pacific Islands (New Zealand); Antarctica
North America; Central America; South America; Eurasia; s Africa; Pacific Islands (New Zealand); Antarctica
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Subspecies varium occurs in a wide range of habitats and displays, accordingly, a wide range of morphological variability. Many features, like the presence of a central strand in the stem, leaf shape, and shape of the leaf apex, are plastic and depend on water availability.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants small to moderately large; stems to 10 cm; costae narrower than 100 µm at base, to mid leaf, percurrent, or rarely short-excurrent; lamina 1-stratose.
subsp. varium
1. Plants extremely large; stems to 15 cm; costae 110-140 µm wide at base, excurrent; lamina sometimes partially 2-stratose, especially at base near costa.
subsp. noterophilum
1. Costae percurrent; alar cells somewhat differentiated; leaves ovate-lanceolate to ovate-triangular; apices obtuse to acute; stem leaves not complanate
H. varium var. varium
1. Costae ending before apex; alar cells not sharply differentiated; leaves broadly ovate; apices acuminate; stem leaves sometimes complanate
H. varium var. humile
Source FNA vol. 28, p. 303. FNA vol. 28, p. 304.
Parent taxa Amblystegiaceae > Hygroamblystegium Amblystegiaceae > Hygroamblystegium > Hygroamblystegium varium
Sibling taxa
H. varium subsp. noterophilum, H. varium var. humile, H. varium var. varium
Subordinate taxa
H. varium subsp. noterophilum, H. varium subsp. varium
H. varium var. humile, H. varium var. varium
Synonyms Leskea varia, Amblystegium varium
Name authority (Hedwig) Mönkemeyer: Hedwigia 50: 275. (1911) unknown
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