Flora of North America species comparison

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Horkelia bolanderi

Bolander's horkelia, border horkelia
Habit Plants openly matted, grayish.
Stems

ascending to erect, 1–3(–4.5) dm, hairs ascending to appressed.
Basal leaves

± planar, 3–8(–9) × 0.7–1.5 cm;

stipules entire;

leaflets 6–9 per side, separate to ± overlapping at least distally, ± cuneate to obovate, 4–10 × 2–7 mm, 1/2 to nearly as wide as long, divided 1/3 to midrib into 3–6 acute, sometimes obtuse, teeth, densely strigulose, often villous marginally or at least apically.
Cauline leaves

(2 or)3–6.
Inflorescences

± congested to open, most flowers arranged in glomerules.
Pedicels

(1–)2–4(–6) mm.
Flowers

10 mm diam.;

epicalyx bractlets elliptic-lanceolate, (1.5–)1.8–3 × 0.5–1 mm, ± 2/3 length of sepals, entire;

hypanthium 1–1.5 × 2–4 mm, less than 1/2 as deep as wide, interior slightly pilose;

sepals spreading to ± reflexed, lanceolate, (2.5–)3–4 mm;

petals oblong to oblanceolate, 3–5.5 × 1.5–2 mm, apex rounded to slightly emarginate;

filaments 0.5–2 × 0.3–0.4 mm, anthers (0.4–)0.5–0.8 mm;

carpels 10–20(–27);

styles (1–)1.5–2 mm.
Achenes

brown to dark brown, 1–1.3 mm, usually faintly roughened.

Horkelia bolanderi
Phenology Flowering summer.
Habitat Edges of vernal lakes, meadows, seeps, in grasslands, conifer woodlands
Elevation 400–1100 m (1300–3600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

W. L. Jepson ([1923–1925]) and P. A. Munz and I. M. Johnston (1925) independently adopted an expanded circumscription of Potentilla bolanderi that included four species of Horkelia recognized by P. A. Rydberg (1908c): H. bernardina, H. bolanderi, H. clevelandii, and H. rydbergii. These were treated as three varieties, which D. D. Keck (1938) converted to subspecies as H. bolanderi subsp. bolanderi, subsp. clevelandii, and subsp. parryi, the last encompassing H. bernardina and H. rydbergii. These subspecies were restored to species rank by B. Ertter (1993c) as H. bolanderi, H. clevelandii, and H. rydbergii, along with the establishment of H. yadonii to accommodate populations from Monterey County assigned to subsp. parryi by Keck.

As now circumscribed, Horkelia bolanderi is known from five locations in the northern Coast Ranges in Colusa, Lake, and Mendocino counties; records from more southerly localities would be referenced to one of the segregate species discussed above.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 9, p. 251.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia
Sibling taxa
H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
Synonyms Potentilla bolanderi
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 338. (1868)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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