Horkelia bolanderi |
Horkelia sericata |
|
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Bolander's horkelia, border horkelia |
Howell's horkelia, silky horkelia |
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Habit | Plants openly matted, grayish. | Plants ± tufted, silvery. |
Stems | ascending to erect, 1–3(–4.5) dm, hairs ascending to appressed. |
ascending to erect, 1.5–4.5 dm, hairs 1 mm proximally, glands absent or sparse distally. |
Basal leaves | ± planar, 3–8(–9) × 0.7–1.5 cm; stipules entire; leaflets 6–9 per side, separate to ± overlapping at least distally, ± cuneate to obovate, 4–10 × 2–7 mm, 1/2 to nearly as wide as long, divided 1/3 to midrib into 3–6 acute, sometimes obtuse, teeth, densely strigulose, often villous marginally or at least apically. |
± cylindric to weakly planar, 3–10 × 0.3–1.2 cm, densely sericeous, often villous on margins apically; stipules usually entire or forked, rarely pinnately divided into linear lobes; leaflets (8–)10–20 per side, ± overlapping, elliptic to flabellate, 2–8 × 1–4 mm, 1/2–2/3 as wide as long, divided ± 1/2+ to midrib into (0–)2–4 elliptic lobes 1–2 mm wide, these not restricted to apex. |
Cauline leaves | (2 or)3–6. |
(2 or)3–5; stipules 3–8 mm, entire or shallowly 1–2-toothed. |
Inflorescences | ± congested to open, most flowers arranged in glomerules. |
open, flowers arranged individually and/or in non-capitate glomerules. |
Pedicels | (1–)2–4(–6) mm. |
1–4 mm. |
Flowers | 10 mm diam.; epicalyx bractlets elliptic-lanceolate, (1.5–)1.8–3 × 0.5–1 mm, ± 2/3 length of sepals, entire; hypanthium 1–1.5 × 2–4 mm, less than 1/2 as deep as wide, interior slightly pilose; sepals spreading to ± reflexed, lanceolate, (2.5–)3–4 mm; petals oblong to oblanceolate, 3–5.5 × 1.5–2 mm, apex rounded to slightly emarginate; filaments 0.5–2 × 0.3–0.4 mm, anthers (0.4–)0.5–0.8 mm; carpels 10–20(–27); styles (1–)1.5–2 mm. |
10 mm diam.; epicalyx bractlets linear-lanceolate, 1.5–3 × 0.5 mm, ± 2/3 length of sepals; hypanthium 1–1.5 × 2–3 mm, ± 1/2 to as deep as wide, interior glabrous; sepals spreading to reflexed, abaxially green to reddish or purplish, 2–4 mm; petals white to pink or red-veined, narrowly obcordate, 3–4.5(–7) × 2–3 mm, apex ± emarginate; filaments 0.5–1.5 × 0.2–0.5 mm, anthers 0.4–0.6 mm; carpels 2–6; styles 1.5–2 mm. |
Achenes | brown to dark brown, 1–1.3 mm, usually faintly roughened. |
brown, 2–2.5 mm, smooth. |
2n | = 28. |
|
Horkelia bolanderi |
Horkelia sericata |
|
Phenology | Flowering summer. | Flowering summer. |
Habitat | Edges of vernal lakes, meadows, seeps, in grasslands, conifer woodlands | Chaparral, oak-conifer woodlands, on serpentine-derived soil |
Elevation | 400–1100 m (1300–3600 ft) | 100–1200 m (300–3900 ft) |
Distribution |
CA
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CA; OR
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Discussion | Of conservation concern. W. L. Jepson ([1923–1925]) and P. A. Munz and I. M. Johnston (1925) independently adopted an expanded circumscription of Potentilla bolanderi that included four species of Horkelia recognized by P. A. Rydberg (1908c): H. bernardina, H. bolanderi, H. clevelandii, and H. rydbergii. These were treated as three varieties, which D. D. Keck (1938) converted to subspecies as H. bolanderi subsp. bolanderi, subsp. clevelandii, and subsp. parryi, the last encompassing H. bernardina and H. rydbergii. These subspecies were restored to species rank by B. Ertter (1993c) as H. bolanderi, H. clevelandii, and H. rydbergii, along with the establishment of H. yadonii to accommodate populations from Monterey County assigned to subsp. parryi by Keck. As now circumscribed, Horkelia bolanderi is known from five locations in the northern Coast Ranges in Colusa, Lake, and Mendocino counties; records from more southerly localities would be referenced to one of the segregate species discussed above. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. D. D. Keck (1938) expanded the circumscription of Horkelia sericata to encompass the plants segregated here as H. howellii, on the grounds that intergradation is too extensive to justify taxonomic recognition of the extremes. In this evaluation, however, most collections can be unequivocally divided between plants with compact silvery-sericeous leaves with entire or forked stipules (H. sericata), and plants with larger, greener leaves and pinnately divided stipules (H. howellii). It is not known if the chromosome count provided by P. A. Munz (1959) was derived from H. howellii or H. sericata. As here circumscribed, Horkelia sericata is a localized taxon known only from Curry County, Oregon, and the Gasquet serpentine area in adjacent Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 251. | FNA vol. 9, p. 265. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae |
Sibling taxa | ||
Synonyms | Potentilla bolanderi | Potentilla sericata |
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 338. (1868) | S. Watson: Proc. Amer. Acad. Arts 20: 364. (1885) |
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