Glyceria fluitans |
Poaceae tribe Meliceae |
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floating mannagrass, floating sweet-grass, glycerie flottante, water manna grass |
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Habit | Plants perennial. | Plants usually perennial, sometimes annual; cespitose, sometimes rhizomatous. | ||||||||||||
Culms | 20-150 cm tall, 2-4 mm thick, erect or spreading, sometimes decumbent and rooting from the lower nodes, distal portion sometimes floating in shallow water. |
annual, not woody, not branching above the base; internodes hollow. |
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Sheaths | glabrous, keeled; ligules 5-15 mm; blades 5-25 cm long, 3-10 mm wide, both surfaces smooth. |
closed for their whole length or almost so; collars without tufts of hair on the sides; auricles sometimes present; ligules hyaline, glabrous, often lacerate, occasionally ciliate, those of the lower and upper cauline leaves usually similar; pseudopetioles absent; blades linear to narrowly lanceolate, venation parallel, cross venation sometimes evident; cross sections non-Kranz, without arm or fusoid cells; epidermes without microhairs, sometimes papillate. |
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Panicles | 10-50 cm long, 2-3 cm wide; branches 3-5 cm, paired or solitary, usually appressed to ascending, divergent at anthesis, with 1-4 spikelets; pedicels 0.8-20 mm. |
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Inflorescences | terminal panicles or racemes; disarticulation above the glumes and beneath the florets or below the glumes. |
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Spikelets | (15)18-39 mm long, 1.7-3.3 mm wide, cylindrical and terete, except slightly laterally compressed at anthesis, rectangular in side view, with 8-16 florets. |
2.5-60 mm, not viviparous, slightly to strongly laterally compressed, with 1-30 florets, proximal florets bisexual, distal 1-3 florets usually sterile, sometimes pistillate, sometimes reduced and amalgamated into a knob- or club-shaped rudiment; rachillas prolonged beyond the base of the distal floret. |
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Glumes | exceeded by the distal florets, shorter than to longer than the adjacent lemmas, mostly membranous, scarious distally, 1-11-veined, apices usually rounded to acute; florets laterally or dorsally compressed; calluses blunt, glabrous or with hairs; lemmas of sexual florets rectangular or ovate, mostly membranous, scarious distally, often with a purplish band adjacent to the scarious apices, (4)5-15-veined, veins not converging distally, often prominent, unawned or awned, awns not branched, apices entire to bilobed or bifid, awns straight, subterminal or from the sinuses; paleas from shorter than to longer than the lemmas, similar in texture, 2-veined, veins keeled, sometimes winged; lodicules 2, fleshy, usually connate into a single structure, without a membranous wing, truncate, not ciliate, not or scarcely veined; anthers 1, 2, or 3; ovaries glabrous; styles 2-branched, bases persistent, branches plumose distally. |
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Lower glumes | 1.3-3.9 mm; upper glumes 2.7-5 mm; rachilla internodes 1.9-2.5 mm; lemmas 5.2-8 mm, midveins extending to within 0.1 mm of the apical margins, scabrous over and between the veins, prickles about 0.05 mm, apices acute, usually entire; paleas from shorter than to 0.6(1.5) mm longer than the lemmas, keels winged, apices bifid, teeth 0.1-0.4 mm, parallel to convergent, sometimes crossing when dry; anthers 1.5-3 mm, usually purple. |
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Caryopses | 2-3 mm. |
ovoid to ellipsoid, longitudinally grooved or not; hila usually linear; embryos less than 1/3 as long as the caryopses. |
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x | = (8)9, 10. |
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2n | = 40. |
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Glyceria fluitans |
Poaceae tribe Meliceae |
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Distribution |
AR; CA; ID; MA; MD; NJ; NY; PA; SD; TN; HI; LB; NS |
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Discussion | Glyceria fluitans is a Eurasian species. In the Americas, it has been collected from British Columbia to California on the west coast, in South Dakota, and from Newfoundland to Pennsylvania on the eastern seaboard. In Europe, it grows in rich, organic, wet soils, often near G. notata, with which it hybridizes. It is less tolerant of trampling than G. notata. Many earlier reports from eastern Canada are based on G. borealis or G. septentrionalis (Dore and McNeill 1980; Scoggan 1978). In western North America, it has been confused with G. xoccidentalis. It tends to differ from all three in its longer lemmas and anthers. Nevertheless, identification of some specimens will prove troublesome. For further discussion, see under the species mentioned. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
There are approximately 130 species and 8 or 9 genera in the Meliceae. Four of the genera are monotypic. Melica and Glyceria, the two largest genera, are well represented in North America. Pleuropogon and Schizachne are primarily North American, but extend into eastern Asia. Molecular studies (e.g., Soreng and Davis 2000; Grass Phylogeny Working Group 2001) show the tribe to be monophyletic and somewhat basal within the Pooideae. Members of the tribe are most easily recognized by the combination of closed leaf sheaths, scarious lemma apices, and non-converging lemma veins. The tribe also differs from other tribes in the Pooideae in having 2 unwinged lodicules that are usually connate into a single structure, and a base chromosome number of 9 or 10. Catabrosa and Briza, whose inclusion in the tribe was suggested by the preliminary results of Mejia-Saules and Bisby (2000), have more membranous lemma margins and free, winged lodicules. Briza also has open leaf sheaths and more convergent lemma veins. Their inclusion is not supported by the molecular data. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 85. | FNA vol. 24, p. 67. | ||||||||||||
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Name authority | (L.) R. Br. | Endl. | ||||||||||||
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