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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower

Jungermann's monkeyflower, liverwort monkey-flower

Habit Perennials, rhizomatous, sometimes rooting at proximal nodes. Perennials, stoloniferous, stolons thin, forming overwintering turions.
Stems

prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident.

decumbent to procumbent, simple or branching near base, 5–38(–60) cm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped, internodes evident.

Leaves

cauline, basal not persistent, often congested;

petiole 0 mm, rarely 1–2(–3) mm;

blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

cauline, basal not persistent;

petiole 2–5(–20) mm;

blade subpalmately to pinnately veined, broadly ovate to broadly lanceolate, 7–35(–40) × 8–25 mm, base rounded, margins sharply, irregularly dentate to denticulate, apex acute to obtuse, surfaces glandular-villous.

Flowers

herkogamous, 4–10, from medial to distal nodes.

herkogamous, 2 or 3, from medial to distal nodes.

Styles

glabrous.

scabrous.

Corollas

yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular;

tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin;

lobe apex rounded.

yellow, with scattered red spots, palate ridges with 2 white patches at tips, bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, (12–)16–20(–24) mm, exserted beyond calyx margin;

limb expanded 8–10 mm, lobes obovate-oblong, apex rounded to truncate.

Fruiting pedicels

(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

15–35 mm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped.

Fruiting calyces

wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate.

plicate-angled, cylindric-urceolate, weakly inflated, 6–12 mm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped, lobes 1–2 mm, apex rounded to mucronate.

Capsules

included, 6–8 mm.

included, 5–9 mm.

Anthers

included, finely hirtellous to hispidulous.

included, glabrous.

2n

= 32.

Erythranthe ptilota

Erythranthe jungermannioides

Phenology Flowering (May–)Jun–Aug. Flowering May–Jul(–Aug).
Habitat Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. Basalt crevices in seepage zones in vertical cliff faces and canyon walls.
Elevation 0–1000(–1900) m. (0–3300(–6200) ft.) 100–400(–1200) m. (300–1300(–3900) ft.)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
from FNA
OR; WA
[WildflowerSearch map]
Discussion

Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves.

A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata.

Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom].

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The occurrence of Erythranthe jungermannioides in the Columbia River Gorge of Klickitat County, Washington (the only record from the state of Washington), is based on an imprecise, unconfirmed observation from the early 1990s (Washington National Heritage Program 2005).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 402. FNA vol. 17, p. 396.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus moschatus var. sessilifolius Mimulus jungermannioides
Name authority G. L. Nesom: Phytoneuron 2017-17: 4. (2017) (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012)
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