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dense boisduvalia, dense spike-primrose, dense willowherb, dense-flower spikeprimrose, dense-flower willowherb, willow herb

Habit Herbs usually with taproot, sometimes with loose network of roots.
Stems

erect or ascending, terete, 4–150 cm, simple or branched with strong central axis, proximal branches ascending or suberect, villous or strigillose, often mixed glandular puberulent distally.

Leaves

opposite and often early-deciduous proximally, alternate and crowded distally, usually subsessile, rarely petiole 1–2 mm, blade usually narrowly lanceolate to sublinear, rarely to lanceolate, 1.4–7.5(–9.2) × 0.5–1.4 cm, base cuneate to attenuate, margins remotely to sharply serrulate, 5–12 teeth per side, lateral veins obscure, 2–5 per side, apex acute, surfaces densely villous and/or strigillose;

bracts broader than cauline leaves, broadly lanceolate to ovate or subrotund, 0.5–2.5 × 0.3–1.8 cm, long-acuminate, sometimes folded on midrib.

Inflorescences

erect spikes, congested, simple, densely villous and strigillose, sometimes mixed glandular puberulent.

Flowers

erect, often hidden within subtending bracts, usually chasmogamous;

buds sessile, narrowly elongate, 2–4 mm;

floral tube 1.3–3.8 × 1–2.2 mm, ring of hairs 0.6–2 mm distal to base inside;

sepals 2–7.5 × 0.5–2.2 mm, apex acute;

petals rose-purple, magenta, pink, or white, 3–9.5(–11.5) × 1.2–5(–6.2) mm, apical notch 0.8–3.8 mm;

filaments dark pink, those of longer stamens 1.5–4.5 mm, those of shorter ones 0.5–1.9 mm;

anthers yellow, 0.5–1.2 × 0.3–0.7 mm;

ovary 2–5 mm, densely villous, often mixed glandular puberulent;

style white, 2.2–5.5(–7.5) mm, glabrous, stigma subcapitate to irregularly 4-lobed, 0.3–0.8 × 0.3–1 mm, surrounded by longer anthers.

Capsules

cylindrical to subfusiform, 4–11 mm, beak to 0.5 mm, central column persistent, surfaces densely villous;

subsessile or pedicel 1–2.5 mm.

Seeds

3–8 per locule, irregularly angular-fusiform, 1.2–1.6(–1.9) × 0.4–1 mm, without a chalazal collar, light brown, surface irregularly reticulate with raised cells.

2n

= 20.

Epilobium densiflorum

Phenology Flowering May–Oct.
Habitat Vernally wet places, moist pastures, woodlands, meadows, along streams and ditches, alluvial valleys, often on low ground in volcanic or sandy soils.
Elevation 0–2600 m. (0–8500 ft.)
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Epilobium densiflorum is an extremely variable species that changes its aspect through the flowering season. Collections made very early in the season may include only well-spaced, narrowly lanceolate leaves, the proximal ones usually opposite, and a short, sparse, somewhat open inflorescence. A late-season collection, even from the same population, may entirely lack cauline leaves, and consist instead of bare, peeling stems topped by dense, tightly imbricate-bracted inflorescences, with each broad bract enclosing a capsule or flower.

Boisduvalia douglasii Spach is an illegitimate substitute for Oenothera densiflora Lindley and pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Pachydium
Sibling taxa
E. anagallidifolium, E. arcticum, E. brachycarpum, E. campestre, E. canum, E. ciliatum, E. clavatum, E. cleistogamum, E. coloratum, E. davuricum, E. densum, E. foliosum, E. glaberrimum, E. hallianum, E. hirsutum, E. hornemannii, E. howellii, E. lactiflorum, E. leptocarpum, E. leptophyllum, E. luteum, E. minutum, E. mirabile, E. montanum, E. nevadense, E. nivium, E. obcordatum, E. obscurum, E. oreganum, E. oregonense, E. pallidum, E. palustre, E. parviflorum, E. rigidum, E. saximontanum, E. septentrionale, E. siskiyouense, E. smithii, E. suffruticosum, E. torreyi
Synonyms Oenothera densiflora, Boisduvalia bipartita, B. densiflora, B. densiflora var. bipartita, B. densiflora var. imbricata, B. densiflora var. montana, B. densiflora var. pallescens, B. densiflora var. salicina, B. imbricata, B. salicina, B. sparsiflora, B. sparsifolia, O. densiflora var. imbricata
Name authority (Lindley) Hoch & P. H. Raven: Phytologia 73: 457. (1993)
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