Epilobium densiflorum |
Epilobium montanum |
|
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dense boisduvalia, dense spike-primrose, dense willowherb, dense-flower spikeprimrose, dense-flower willowherb, willow herb |
broad-leaf willow-herb, tooth willow-herb, willowherb |
|
Habit | Herbs usually with taproot, sometimes with loose network of roots. | Herbs with short, basal stolons terminating in fleshy or leafy rosettes at, or just below, ground level. |
Stems | erect or ascending, terete, 4–150 cm, simple or branched with strong central axis, proximal branches ascending or suberect, villous or strigillose, often mixed glandular puberulent distally. |
erect, terete, (5–)20–95 cm, often well branched, especially distally, densely strigillose proximal to inflorescence, without decurrent lines, mixed strigillose and glandular puberulent distally. |
Leaves | opposite and often early-deciduous proximally, alternate and crowded distally, usually subsessile, rarely petiole 1–2 mm, blade usually narrowly lanceolate to sublinear, rarely to lanceolate, 1.4–7.5(–9.2) × 0.5–1.4 cm, base cuneate to attenuate, margins remotely to sharply serrulate, 5–12 teeth per side, lateral veins obscure, 2–5 per side, apex acute, surfaces densely villous and/or strigillose; bracts broader than cauline leaves, broadly lanceolate to ovate or subrotund, 0.5–2.5 × 0.3–1.8 cm, long-acuminate, sometimes folded on midrib. |
opposite proximal to inflorescence, alternate distally, petiole 1–6 mm; blade dark green, narrowly ovate to ovate, 3–8 × 1.5–3.4 cm, subequal to internodes, base broadly cuneate to truncate, margins irregularly serrulate with 20–30 teeth per side, veins prominent, 3–5 per side, apex acuminate to acute, surfaces strigillose, especially along veins and margins; bracts much reduced. |
Inflorescences | erect spikes, congested, simple, densely villous and strigillose, sometimes mixed glandular puberulent. |
nodding to erect, racemes or loose panicles, strigillose and glandular puberulent. |
Flowers | erect, often hidden within subtending bracts, usually chasmogamous; buds sessile, narrowly elongate, 2–4 mm; floral tube 1.3–3.8 × 1–2.2 mm, ring of hairs 0.6–2 mm distal to base inside; sepals 2–7.5 × 0.5–2.2 mm, apex acute; petals rose-purple, magenta, pink, or white, 3–9.5(–11.5) × 1.2–5(–6.2) mm, apical notch 0.8–3.8 mm; filaments dark pink, those of longer stamens 1.5–4.5 mm, those of shorter ones 0.5–1.9 mm; anthers yellow, 0.5–1.2 × 0.3–0.7 mm; ovary 2–5 mm, densely villous, often mixed glandular puberulent; style white, 2.2–5.5(–7.5) mm, glabrous, stigma subcapitate to irregularly 4-lobed, 0.3–0.8 × 0.3–1 mm, surrounded by longer anthers. |
erect; buds 4–5x 1.5–3 mm; pedicel 2–18 mm; floral tube 1.2–2 × 1.5–2.5 mm, conspicuous ring of spreading hairs at mouth inside; sepals often flushed red, keeled, 5–6.5 × 1.2–1.7 mm, abaxial surface strigillose and glandular puberulent; petals rose-purple, obcordate, 7.5–10 × 4–5.5 mm, apical notch 2.7–4.2 mm; filaments light pink, those of longer stamens 3.5–5.5 mm, those of shorter ones 1.5–2.4 mm; anthers yellow, 1–1.2 × 0.5–0.6 mm; ovary 20–30 mm, strigillose and glandular puberulent; style rose-purple to white, 3.5–7.5 mm, glabrous or with scattered long hairs near base, stigma deeply 4-lobed, 1.5–2 × 2.5–4 mm, lobes 1.2–1.8 mm, exserted beyond or, sometimes, surrounded by anthers. |
Capsules | cylindrical to subfusiform, 4–11 mm, beak to 0.5 mm, central column persistent, surfaces densely villous; subsessile or pedicel 1–2.5 mm. |
40–80 mm, surfaces strigillose and glandular puberulent; pedicel 8–20 mm. |
Seeds | 3–8 per locule, irregularly angular-fusiform, 1.2–1.6(–1.9) × 0.4–1 mm, without a chalazal collar, light brown, surface irregularly reticulate with raised cells. |
obovoid, 1–1.2 × 0.4–0.5 mm, chalazal collar inconspicuous, 0.6–0.8 mm wide, brown, surface coarsely papillose; coma readily detached, dull white, 5–6 mm. |
2n | = 20. |
= 36. |
Epilobium densiflorum |
Epilobium montanum |
|
Phenology | Flowering May–Oct. | Flowering Jul–Aug. |
Habitat | Vernally wet places, moist pastures, woodlands, meadows, along streams and ditches, alluvial valleys, often on low ground in volcanic or sandy soils. | Moist rocky banks, gravelly slopes, open woods, disturbed or waste areas. |
Elevation | 0–2600 m. (0–8500 ft.) | 0–150[–1500] m. (0–500[–4900] ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; BC; Mexico (Baja California)
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ME; BC; NF; Europe; Asia [Introduced in North America; introduced also in Pacific Islands (New Zealand)] |
Discussion | Epilobium densiflorum is an extremely variable species that changes its aspect through the flowering season. Collections made very early in the season may include only well-spaced, narrowly lanceolate leaves, the proximal ones usually opposite, and a short, sparse, somewhat open inflorescence. A late-season collection, even from the same population, may entirely lack cauline leaves, and consist instead of bare, peeling stems topped by dense, tightly imbricate-bracted inflorescences, with each broad bract enclosing a capsule or flower. Boisduvalia douglasii Spach is an illegitimate substitute for Oenothera densiflora Lindley and pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Epilobium montanum is native and widely distributed throughout Europe and in Asia at least as far east as the Ural Mountains and the Caucasus, and in Japan (A. W. Lievens and P. C. Hoch 1999). It also is questionably naturalized on Campbell Island, New Zealand (P. H. Raven and T. E. Raven 1976). Although E. montanum was first collected in 1894 in the vicinity of St. John’s, Newfoundland, and clearly naturalized in that area, it does not appear to have spread appreciably in eastern North America, unlike European adventives such as E. hirsutum or E. parviflorum. The collection by F. C. Seymour in 1971 in Maine may be an independent introduction. Collections of E. montanum in the University of British Columbia Botanical Garden in Vancouver appeared to have been ephemeral, but more recent collections in ruderal areas in that region suggest that it has naturalized there. Like other European introductions, E. montanum has the BB chromosome arrangement (S. R. Seavey and P. H. Raven 1977). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Pachydium | Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Epilobium |
Sibling taxa | ||
Synonyms | Oenothera densiflora, Boisduvalia bipartita, B. densiflora, B. densiflora var. bipartita, B. densiflora var. imbricata, B. densiflora var. montana, B. densiflora var. pallescens, B. densiflora var. salicina, B. imbricata, B. salicina, B. sparsiflora, B. sparsifolia, O. densiflora var. imbricata | Chamaenerion montanum |
Name authority | (Lindley) Hoch & P. H. Raven: Phytologia 73: 457. (1993) | Linnaeus: Sp. Pl. 1: 348. (1753) |
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