Eleocharis engelmannii |
Eleocharis obtusa |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
blunt spike-rush, blunt spikesedge, broad spike spikerush, egg-shape spike-rush, éléocharide obtuse |
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Culms | 2–40 cm × 0.5–1.5(–2) mm. |
3–50(–90) cm × 0.2–2 mm. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
broadly ovoid (to ellipsoid or lanceoloid), apex rounded (to acute), (2–)5–13 × (2–)3–4 mm; floral scales 15–150+, 8–20 per mm of rachilla, orange-brown (to stramineous), elliptic, 1.5–2.5 × 1–1.5 mm, midribs seldom keeled, apex broadly rounded. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles (5–)6–7, rarely 0, brown, stout, slightly to usually greatly exceeding tubercle; stamens usually 3; anthers brown to yellow, 0.3–0.6 mm; styles usually 3-fid and 2-fid in same spikelet. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
0.9–1.2(–1.3) × 0.7–0.9 mm. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
deltoid 0.35–0.5 × (0.4–)0.5–0.8 mm, 1/3–2/3 as high as wide, 1/3–1/2 as high and 2/3–9/10 as wide as achene. |
2n | = 10. |
= 10. |
Eleocharis engelmannii |
Eleocharis obtusa |
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Phenology | Fruiting spring–fall. | Fruiting summer–fall. |
Habitat | Fresh shores, marshes, disturbed places | Fresh shores, marshes, disturbed places |
Elevation | 30–2400 m (100–7900 ft) | 10–1600 m (0–5200 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; RI; SC; TN; TX; VA; VT; WA; WI; WV; WY; HI; BC; NS; ON; PE; QC
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Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Extremely uncommon plants of Eleocharis obtusa without perianth bristles may be called E. obtusa var. peasei (type from New Hampshire). Robust plants with distinct caudices, floral scales 2.5 mm, and achenes 1.2–1.3 mm (Eleocharis obtusa var. gigantea Fernald) are rare (specimens seen from the Washington-British Columbia border [type], Arkansas, and the Hawaiian Islands). Dwarf plants (E. obtusa var. jejuna Fernald, type from Maine), with unusually small achenes and floral scales, and tubercles often less than 0.5 mm wide, are occasional in the East and are easily confused with E. ovata and E. aestuum. A few specimens are intermediate with E. engelmannii. Eleocharis obtusa is sometimes treated as conspecific with E. ovata, which consistently differs in its mostly 2-fid styles, mostly two stamens, and especially its narrower tubercles (B. M. H. Larson and P. M. Catling 1996). Eleocharis macounii Fernald has been treated as a synonym of E. obutsa (H. K. Svenson 1957) but is more probably a hybrid between E. intermedia and E. obtusa (P. M. Catling and S. G. Hay 1993; see 34. E. intermedia). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 105. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | Scirpus obtusus, E. obtusa var. ellipsoidales, E. obtusa var. gigantea, E. obtusa var. jejuna, E. obtusa var. peasei |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | (Willdenow) Schultes: Mant. 2: 89. (1824) |
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