Eleocharis engelmannii |
Eleocharis erythropoda |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
bald spike-rush, red-footed spikesedge, red-stem spikerush, redfoot spike-rush, spikesedge, éléocharide à tiges rouges |
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Habit | Plants perennial, mat-forming; rhizomes evident, long, 0.5–1.5 mm thick, soft to firm, cortex often breaking loose, longer internodes 5–30 mm, scales often fugaceous, 4–8 mm, membranous, not fibrous. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
terete, often with 8–12 blunt ridges when dry, 8–80 cm × 0.3–1.4 mm, firm to soft, internally spongy. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths persistent, not splitting, proximally red (to stramineous), distally green to stramineous, usually inflated, often callose, membranous to papery, apex often red-brown, broadly obtuse to subacute, tooth sometimes present, to 0.1 mm. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
ovoid to lanceoloid or nearly cylindric, 3–18 × 2–3(–4) mm, apex acute (to obtuse); proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales often spreading in fruit, 15–50, 4–5 per mm of rachilla, medium brown to sometimes red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, apex rounded, in distal part lanceolate, apex entire, acute, 2–3.5 × 1.5–1.7 mm, mostly carinate. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles 4 or absent, light brown to stramineous, stout, usually equal, equaling achene to slightly exceeding tubercle; stamens 3; anthers dark yellow to stramineous, 1–1.8 mm, apiculate; styles 2-fid. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
not persistent, dark yellow, stramineous, or dark brown, obovoid to obpyriform, biconvex, angles obscure, 0.9–1.6 × 0.7–1.2 mm, apex rounded, neck absent to long, smooth at 30X, or sometimes finely rugulose at 10–30X with 20 or more horizontal ridges in a vertical series. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
brown to whitish, pyramidal, much higher than wide to, lower than wide, 0.35–0.65 × 0.2–0.6 mm. |
2n | = 10. |
= 16, 18, 19, 20. |
Eleocharis engelmannii |
Eleocharis erythropoda |
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Phenology | Fruiting spring–fall. | Fruiting summer. |
Habitat | Fresh shores, marshes, disturbed places | Non-calcareous or calcareous fresh or brackish shores, marshes, meadows, fens, disturbed places |
Elevation | 30–2400 m (100–7900 ft) | 0–2300 m (0–7500 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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AK; AR; AZ; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
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Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis erythropoda is extremely variable. Intermediates with E. palustris variant b are common in the region of sympatry in the East, and with E. macrostachya variant b in the West, and E. erythropoda is sometimes difficult to distinguish from E. uniglumis. Eleocharis calva Torrey is an invalid name. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 75. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | Scirpus glaucus |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | Steudel: Syn. Pl. Glumac. 2: 76. (1855) |
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