Dermatophyllum
Dermatophyllum arizonicum
mescal bean
Arizona necklacepod
erect, twigs densely pubescent or glabrescent.
alternate, odd-pinnate;
stipules present, caducous, linear to deltate;
petiolate, petiole 1–1.5 cm;
leaflets 5–13[–17], alternate to subopposite, stipels minute or absent, linear, blade leathery, margins entire, thickened, surfaces pubescent or glabrescent.
rachis 3–7 cm;
leaflets 5–11, blades lanceolate to elliptic, 1–2.5(–4) × 0.4–1.3 cm, base cuneate to rounded, apex acute, apiculate.
2–8-flowered, congested, 2.5–4 cm;
bracts lanceolate, apex acute.
2–15[–75]-flowered, terminal or axillary, racemes;
bracts present, caducous;
bracteoles persistent or caducous, 2.
2–5 mm.
papilionaceous;
calyx campanulate (sometimes gibbous), lobes 5, sometimes connate adaxially;
corolla usually purple, blue-purple, lilac, or lavender [violet], rarely white, glabrous;
keel petals usually partly connate;
stamens 10, distinct or connate proximally;
anthers dorsifixed.
ascending, 16–24 mm;
calyx obconic, 10–11(–15) mm;
corolla usually purple or lilac, rarely white.
legumes, stipitate, torose to torulose, straight to slightly curved, compressed and oblong, or subglobose to cylindrical, indehiscent, papery, leathery, or woody, appressed-pubescent.
tan, torose, straight to slightly curved, compressed, oblong, 5–10(–12) × 1–1.4 cm, papery to almost leathery.
1–10, usually red or dull red to reddish brown, rarely orange or yellow, reniform to subglobose, margins angular.
3–7(–10), dull red to reddish brown, 7–11 mm.
= 9.
= 18.
Dermatophyllum
Dermatophyllum arizonicum
Species 6 (3 in the flora).
Two other generic names have been used for taxa now placed in Dermatophyllum, in addition to their placement in Sophora: Agastianis Rafinesque and Calia Terán & Berlandier (G. P. Yakovlev 1968). Agastianis is superfluous and an illegitimate substitute for the nomenclaturally rejected name Broussonetia Ortega (1798). Calia is considered a later homonym of Calea Linnaeus (1763, Asteraceae; K. N. Gandhi et al. 2011).
Dermatophyllum is a segregate from Sophora. It is considered distinct from Sophora by its woody habit; thick, leathery leaflets; blue, violet, or white petals; calyx with obvious teeth or lobes; flattened to terete legumes; and geographic range. The distinction is supported by molecular data indicating that Dermatophyllum (as Calia) falls into a separate clade from Styphnolobium and other Sophora species (K. N. Gandhi et al. 2011; G. P. Lewis et al. 2005; R. T. Pennington et al. 2001; M. F. Wojciechowski et al. 2004).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Dermatophyllum arizonicum is known from Graham, Mohave, and Yavapai counties. This species, D. guadalupense, and the Mexican D. gypsophilum (B. L. Turner & A. M. Powell) Vincent, D. juanhintonianum (B. L. Turner) B. L. Turner, and D. purpusii (Brandegee) Vincent, form a closely related group (M. Izaddoost 1975; D. K. Northington 1976; V. E. Rudd 1972; B. L. Turner 2012). D. Isely (1981) suggested that some of these taxa may not deserve recognition as distinct species.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaflet blades (2–)2.5–5(–8) cm; pedicels 10–15 mm; legumes woody, subglobose to cylindrical, torulose. | D. secundiflorum |
1. Leaflet blades 1–2.5(–4) cm; pedicels 2–5 mm; legumes papery to almost leathery, compressed and oblong, torose. | → 2 |
2. Leaflets 5–11, blades lanceolate to elliptic, apices acute, apiculate; floral bracts lance- olate, apices acute. | D. arizonicum |
2. Leaflets 9–13, blades elliptic to ovate, apices rounded, often emarginate, rarely apiculate; floral bracts lanceolate to elliptic, apices acute to rounded, abruptly apiculate. | D. guadalupense |
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