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mescal bean

Habit Shrubs or trees, unarmed.
Stems

erect, twigs densely pubescent or glabrescent.

Leaves

alternate, odd-pinnate;

stipules present, caducous, linear to deltate;

petiolate, petiole 1–1.5 cm;

leaflets 5–13[–17], alternate to subopposite, stipels minute or absent, linear, blade leathery, margins entire, thickened, surfaces pubescent or glabrescent.

Inflorescences

2–15[–75]-flowered, terminal or axillary, racemes;

bracts present, caducous;

bracteoles persistent or caducous, 2.

Flowers

papilionaceous;

calyx campanulate (sometimes gibbous), lobes 5, sometimes connate adaxially;

corolla usually purple, blue-purple, lilac, or lavender [violet], rarely white, glabrous;

keel petals usually partly connate;

stamens 10, distinct or connate proximally;

anthers dorsifixed.

Fruits

legumes, stipitate, torose to torulose, straight to slightly curved, compressed and oblong, or subglobose to cylindrical, indehiscent, papery, leathery, or woody, appressed-pubescent.

Seeds

1–10, usually red or dull red to reddish brown, rarely orange or yellow, reniform to subglobose, margins angular.

x

= 9.

Dermatophyllum

Distribution
sw United States; sc United States; Mexico
[BONAP county map]
Discussion

Species 6 (3 in the flora).

Two other generic names have been used for taxa now placed in Dermatophyllum, in addition to their placement in Sophora: Agastianis Rafinesque and Calia Terán & Berlandier (G. P. Yakovlev 1968). Agastianis is superfluous and an illegitimate substitute for the nomenclaturally rejected name Broussonetia Ortega (1798). Calia is considered a later homonym of Calea Linnaeus (1763, Asteraceae; K. N. Gandhi et al. 2011).

Dermatophyllum is a segregate from Sophora. It is considered distinct from Sophora by its woody habit; thick, leathery leaflets; blue, violet, or white petals; calyx with obvious teeth or lobes; flattened to terete legumes; and geographic range. The distinction is supported by molecular data indicating that Dermatophyllum (as Calia) falls into a separate clade from Styphnolobium and other Sophora species (K. N. Gandhi et al. 2011; G. P. Lewis et al. 2005; R. T. Pennington et al. 2001; M. F. Wojciechowski et al. 2004).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflet blades (2–)2.5–5(–8) cm; pedicels 10–15 mm; legumes woody, subglobose to cylin­drical, torulose.
D. secundiflorum
1. Leaflet blades 1–2.5(–4) cm; pedicels 2–5 mm; legumes papery to almost leathery, compressed and oblong, torose.
→ 2
2. Leaflets 5–11, blades lanceolate to elliptic, apices acute, apiculate; floral bracts lance- olate, apices acute.
D. arizonicum
2. Leaflets 9–13, blades elliptic to ovate, apices rounded, often emarginate, rarely apiculate; floral bracts lanceolate to elliptic, apices acute to rounded, abruptly apiculate.
D. guadalupense
Source FNA vol. 11. Authors: Michael A. Vincent, Denis M. Kearns.
Parent taxa Fabaceae > subfam. Faboideae
Subordinate taxa
D. arizonicum, D. guadalupense, D. secundiflorum
Name authority Scheele: Linnaea 21: 458. (1848)
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