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Bermuda grass, chiendent pied-de-poule, common bermudagrass, devilgrass, grama-seda, manienie, motie molulu

Bermuda grass, cynodon, dog's-tooth grass

Habit Plants stoloniferous, usually also rhizomatous. Plants perennial; sometimes stoloniferous, sometimes also rhizomatous, often forming dense turf.
Culms

5-40(50) cm, not becoming woody.

4-100 cm.

Sheaths

glabrous or with scattered hairs;

collars usually with long hairs, particularly at the margins;

ligules about 0.5 mm, of hairs;

blades 1-6(16) cm long, (1)2-4(5) mm wide, flat at maturity, conduplicate or convolute in bud, glabrous or the adaxial surfaces pilose.

open;

auricles absent;

ligules of hairs or membranous;

blades flat, conduplicate, convolute, or involute, sometimes disarticulating.

Panicles

with (2)4-6(9) branches;

branches 2-6 cm, in a single whorl, axes triquetrous.

Inflorescences

terminal, digitate or subdigitate panicles of spikelike branches;

branches (1)2-20, 1-sided, with 2 rows of solitary, subsessile, appressed, imbricate spikelets.

Spikelets

2-3.2 mm.

laterally compressed, with 1(-3) florets, only the lowest floret functional;

rachilla extension usually present, sometimes terminating in a reduced floret;

disarticulation above the glumes.

Glumes

usually shorter than the lemmas, membranous, keeled, usually muticous;

lower glumes 1-veined;

upper glumes 1-3-veined, occasionally shortly awned;

lemmas membranous to cartilaginous, 3-veined, keeled, keels with hairs, occasionally winged, apices mucronate or muticous;

paleas about as long as the lemmas, 2-keeled;

anthers 3;

style branches 2, plumose;

lodicules 2.

Lower

glumes 1.5-2 mm;

upper glumes 1.4-2.3 mm;

lemmas 1.9-3.1 mm, keels not winged, pubescent, margins usually less densely pubescent;

anthers dehiscent at maturity;

paleas glabrous.

x

= 9.

2n

= 18, 36.

Cynodon dactylon

Cynodon

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; HI; PR; BC; Virgin Islands
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; HI; PR; BC; Virgin Islands
[BONAP county map]
Discussion

Cynodon dactylon is a variable species, but taxonomists disagree on just how variable. Caro and Sanchez (1969) limited C. dactylon to plants with conduplicate leaves, placing those with convolute leaves in a number of other species, such as C. affinis Caro & Sanchez and C. aristiglumis Caro & Sanchez; de Wet and Harlan (1970) do not mention this character in their study of Cynodon, Caro and Sanchez also employed several other characters in the key separating C. dactylon from the species with convolute immature leaves, but the overlap between the two sides of the lead is substantial. Pending further study, the broader interpretation, in which C. dactylon includes plants with both convolute and conduplicate leaves, has been adopted.

Several varieties of C. dactylon have been described, in addition to which numerous cultivars have been developed, some as turf grasses for lawns or putting greens, others as pasture or forage grasses. Their useful range is limited because C. dactylon is not cold hardy, going dormant and turning brown when nighttime temperatures fall below freezing or average daytime temperatures are below 10°C.

The most commonly encountered variety, both in the Flora region and in other parts of the world, is C. dactylon var. dactylon, largely because it thrives in severely disturbed, exposed sites; it does not invade natural grass-lands or forests. Determining how many other varieties are established in the Flora region is almost impossible, because there has been no global study of variation in the species. The presence of numerous cultivars complicates an already difficult problem. The two varieties keyed out below are the only two that grow in the Flora region according to de Wet and Harlan (1970), but these authors do not appear to have considered the taxa recognized by Caro and Sanchez (1969). For most purposes, it is probably neither necessary nor feasible to identify the variety of C. dactylon encountered.

Cynodon dactylon is considered a weed in many countries and it is true that, once established, it is difficult to eradicate. It does, however, have some redeeming values. It is rich in vitamin C, and its leaves are sometimes used for an herbal tea. It is claimed to have various medicinal properties, but these have not been verified. It is considered a good pasture grass, in addition to which it is sometimes grown as an ornamental and for erosion control on exposed soils.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cynodon is a genus of nine species, all of which are native to tropical regions of the Eastern Hemisphere. Several species are used as lawn and forage grasses in tropical and warm-temperate regions. The most widespread species, Cynodon dactylon, is also the most frequently encountered species in the Flora region. It is used for lawns, putting greens, and pastures in southern portions of the region, but is generally considered a weed in other parts.

The status of several species in the Flora region is unclear. Species other than C. dactylon usually grow only under cultivation, but there are scattered records of populations of other species from the southern United States that appear to have become established. Cultivars of C. aethiopicus and C. nlemfuënsis are used for pasture primarily in tropical Florida. Cynodon transvaalensis has had limited commercial distribution as a turf grass.

Many cultivars of Cynodon have been developed, some from hybrids between it and other species such as C. transvaalensis, C. aethiopicus, and C. nlemfuënsis. The cultivars may exhibit combinations of features that are not found in the wild species, making it difficult to accommodate them in a key.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Rhizomes near the surface (sometimes surfacing for a short distance before submerging again), the tips eventually surfacing and, like the lateral buds, producing tillers
var. dactylon
1. Rhizomes growing up to 1 m deep, the tips remaining below ground, only the lateral buds producing tillers
var. aridus
1. Lemma keels winged; panicle branches with flattened axes (subg. Pterolemma)
C. incompletus
1. Lemma keels not winged; panicle branches with triquetrous axes (subg. Cynodon).
→ 2
2. Glumes 0.1-0.6 mm long
C. plectostachyus
2. Glumes 1.1-2.6 mm long.
→ 3
3. Panicles with 1-3(4) branches; culms 5-30 cm tall; blades 1-1.5 mm wide
C. transvaalensis
3. Panicles with (2)4-20 branches; culms 5-100 cm tall; blades (1)2-7 mm wide.
→ 4
4. Panicles with 2-6(9) branches in a single whorl; culms 5-40(50) cm tall.
→ 5
5. Panicles with (2)4-6(9) branches; anthers dehiscent at maturity
C. dactylon
5. Panicles with 2-4 branches; anthers indehiscent at maturity
C. ×magennisii
4. Panicles with 4-20 branches in 1-5 whorls; culms 20-100 cm tall.
→ 6
6. Lemma keels glabrous or with a few scattered hairs; panicle branches usually in 2-5 whorls, stiff, frequently red or purple; culms 25-100 cm tall, woody
C. aethiopicus
6. Lemma keels shortly pubescent; panicle branches usually in 1 whorl, lax, usually green; culms 20-60 cm tall, not woody
C. nlemfuënsis
Source FNA vol. 25, p. 238. FNA vol. 25, p. 235. Author: Mary E. Barkworth;.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Cynodon Poaceae > subfam. Chloridoideae > tribe Cynodonteae
Sibling taxa
C. aethiopicus, C. incompletus, C. nlemfuënsis, C. plectostachyus, C. transvaalensis, C. ×magennisii
Subordinate taxa
C. dactylon var. aridus, C. dactylon var. dactylon
C. aethiopicus, C. dactylon, C. incompletus, C. nlemfuënsis, C. plectostachyus, C. transvaalensis, C. ×magennisii
Synonyms Cynosurus dactylon, C. erectus, Capriola dactylon
Name authority (L.) Pers. Rich.
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