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brownhair lacefern


scales brown, linear, 10–20 × 0.8–1.5 mm, lax, not densely tangled or woollike.


1-pinnate-pinnatifid, glabrous or pubescent on both surfaces, glandular abaxially and occasionally adaxially, glands pale yellow, ca. 0.5 mm.

Ultimate segments

4–7 mm wide, margins ciliate.


6–10(–15) pairs per segment, unbranched.


medial to supramedial;

indusia present but soon deciduous or completely absent.


pinnae 8–18 × 2–3 cm, equilateral, incised more than 3/4 distance to costae.

Ctenitis submarginalis

Habitat Cypress swamps, hammocks, old forested spoil banks
Elevation 0 m (0 ft)
from FNA
FL; LA; e Mexico; s Mexico; Central America; West Indies in Hispaniola; South America to Uruguay
[BONAP county map]

Combining authorship of the accepted name sometimes has been incorrectly attributed to E. B. Copeland (1947). The Louisiana population of Ctenitis submarginalis occurs more than 960 km from populations in Florida and represents the northernmost locality for the species (G. P. Landry and W. D. Reese 1991). Unlike most ferns in North America, the plants in the Louisiana population are nonseasonal, producing leaves and sori throughout the year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2.
Parent taxa Dryopteridaceae > Ctenitis
Sibling taxa
C. sloanei
Synonyms Polypodium submarginale, Dryopteris submarginalis
Name authority (Langsdorff & Fischer) Ching: Sunyatsenia 5: 250. (1940)
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