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brownhair lacefern

Florida lacefern, Florida tree fern, red-hair comb fern


scales brown, linear, 10–20 × 0.8–1.5 mm, lax, not densely tangled or woollike.

scales orangish, rarely brown, linear or filiform, 20–40 × 0.4–1.5 mm, densely tangled and woollike.


1-pinnate-pinnatifid, glabrous or pubescent on both surfaces, glandular abaxially and occasionally adaxially, glands pale yellow, ca. 0.5 mm.

2–4-pinnate-pinnatifid, glabrous or glandular on both surfaces;

glands pale yellow, ca. 0.1 mm, appressed.

Ultimate segments

4–7 mm wide, margins ciliate.

2–4 mm wide;

margins ciliate.


6–10(–15) pairs per segment, unbranched.

3–6 pairs per segment, unbranched or 1-forked.


medial to supramedial;

indusia present but soon deciduous or completely absent.

medial to inframedial;

indusia present but soon deciduous and therefore appearing absent.


pinnae 8–18 × 2–3 cm, equilateral, incised more than 3/4 distance to costae.

pinnae 22–50 × 10–23 cm, inequilateral, elongate basiscopically.


= 82.

Ctenitis submarginalis

Ctenitis sloanei

Habitat Cypress swamps, hammocks, old forested spoil banks Wooded limestone ledges, hammocks, cypress swamps
Elevation 0 m (0 ft) 0 m (0 ft)
from FNA
FL; LA; e Mexico; s Mexico; Central America; West Indies in Hispaniola; South America to Uruguay
[BONAP county map]
from FNA
FL; s Mexico; Central America; West Indies in Antilles; Trinidad; South America in Colombia; Ecuador; Peru; and Venezuela
[BONAP county map]

Combining authorship of the accepted name sometimes has been incorrectly attributed to E. B. Copeland (1947). The Louisiana population of Ctenitis submarginalis occurs more than 960 km from populations in Florida and represents the northernmost locality for the species (G. P. Landry and W. D. Reese 1991). Unlike most ferns in North America, the plants in the Louisiana population are nonseasonal, producing leaves and sori throughout the year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The names Ctenitis ampla (Humboldt & Bonpland ex Willdenow) Ching and Dryopteris ampla (Humboldt & Bonpland ex Willdenow) Kuntze have been misapplied to this taxon. Ctenitis sloanei and C. submarginalis both have numerous scales at the base of the petiole; in C. sloanei, however, the scales form a large, conspicuous, tangled tuft.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Ctenitis Dryopteridaceae > Ctenitis
Sibling taxa
C. sloanei
C. submarginalis
Synonyms Polypodium submarginale, Dryopteris submarginalis Polypodium sloanei
Name authority (Langsdorff & Fischer) Ching: Sunyatsenia 5: 250. (1940) (Poeppig ex Sprengel) C. V. Morton: Amer. Fern J. 59: 66. (1969)
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