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comb fern, lacefern

brownhair lacefern

Habit Plants generally terrestrial.
Stems

erect to obliquely ascending, stolons absent.

Leaves

monomorphic, evergreen or dying back in winter.

Petiole

2/3 to equaling length of blade, base not swollen;

vascular bundles more than 3, arranged in an arc, ± round in cross section.

scales brown, linear, 10–20 × 0.8–1.5 mm, lax, not densely tangled or woollike.

Blade

lanceolate to deltate, 1–4-pinnate-pinnatifid, gradually reduced distally to confluent, pinnatifid apex, herbaceous.

1-pinnate-pinnatifid, glabrous or pubescent on both surfaces, glandular abaxially and occasionally adaxially, glands pale yellow, ca. 0.5 mm.

Ultimate segments

4–7 mm wide, margins ciliate.

Pinnae

not articulate to rachis, segment margins nearly entire to crenulate, ciliate;

proximal pinnae not reduced, sometimes basal pair much the longest, sessile to petiolulate, equilateral or inequilateral with basiscopic side more developed (pinnules noticeably longer);

costae adaxially rounded or flat, not grooved;

indument of linear to lanceolate scales and often multicellular glandular hairs abaxially, of multicellular reddish hairs adaxially.

Veins

free, simple or forked.

6–10(–15) pairs per segment, unbranched.

Sori

in 1 row between midrib and margin, round;

indusia round-reniform, attached at narrow sinus, sometimes small or seemingly absent, persistent or caducous.

medial to supramedial;

indusia present but soon deciduous or completely absent.

Spores

brownish, usually spiny, sometimes prominently cristate, rarely finely reticulate.

Basal

pinnae 8–18 × 2–3 cm, equilateral, incised more than 3/4 distance to costae.

x

= 41.

Ctenitis

Ctenitis submarginalis

Habitat Cypress swamps, hammocks, old forested spoil banks
Elevation 0 m (0 ft)
Distribution
from USDA
Nearly worldwide in the tropics
[BONAP county map]
from FNA
FL; LA; e Mexico; s Mexico; Central America; West Indies in Hispaniola; South America to Uruguay
[BONAP county map]
Discussion

Species ca. 100 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Combining authorship of the accepted name sometimes has been incorrectly attributed to E. B. Copeland (1947). The Louisiana population of Ctenitis submarginalis occurs more than 960 km from populations in Florida and represents the northernmost locality for the species (G. P. Landry and W. D. Reese 1991). Unlike most ferns in North America, the plants in the Louisiana population are nonseasonal, producing leaves and sori throughout the year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Blades ovate-lanceolate, 2-4-pinnate-pinnatifid; basal pinnae inequilateral, elongate basiscopically.
C. sloanei
1. Blades oblong or narrowly lanceolate, 1-pinnate-pinnatifid; basal pinnae equilateral.
C. submarginalis
Source FNA vol. 2. Author: Robbin C. Moran. FNA vol. 2.
Parent taxa Dryopteridaceae Dryopteridaceae > Ctenitis
Sibling taxa
C. sloanei
Subordinate taxa
C. sloanei, C. submarginalis
Synonyms Dryopteris subg. C. Polypodium submarginale, Dryopteris submarginalis
Name authority (C. Christensen) C. Christensen: in Verdoorn et al., Man. Pteridol. 543. (1938) (Langsdorff & Fischer) Ching: Sunyatsenia 5: 250. (1940)
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