Ctenitis |
Ctenitis submarginalis |
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comb fern, lacefern |
brownhair lacefern |
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Habit | Plants generally terrestrial. | |||||
Stems | erect to obliquely ascending, stolons absent. |
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Leaves | monomorphic, evergreen or dying back in winter. |
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Petiole | 2/3 to equaling length of blade, base not swollen; vascular bundles more than 3, arranged in an arc, ± round in cross section. |
scales brown, linear, 10–20 × 0.8–1.5 mm, lax, not densely tangled or woollike. |
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Blade | lanceolate to deltate, 1–4-pinnate-pinnatifid, gradually reduced distally to confluent, pinnatifid apex, herbaceous. |
1-pinnate-pinnatifid, glabrous or pubescent on both surfaces, glandular abaxially and occasionally adaxially, glands pale yellow, ca. 0.5 mm. |
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Ultimate segments | 4–7 mm wide, margins ciliate. |
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Pinnae | not articulate to rachis, segment margins nearly entire to crenulate, ciliate; proximal pinnae not reduced, sometimes basal pair much the longest, sessile to petiolulate, equilateral or inequilateral with basiscopic side more developed (pinnules noticeably longer); costae adaxially rounded or flat, not grooved; indument of linear to lanceolate scales and often multicellular glandular hairs abaxially, of multicellular reddish hairs adaxially. |
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Veins | free, simple or forked. |
6–10(–15) pairs per segment, unbranched. |
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Sori | in 1 row between midrib and margin, round; indusia round-reniform, attached at narrow sinus, sometimes small or seemingly absent, persistent or caducous. |
medial to supramedial; indusia present but soon deciduous or completely absent. |
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Spores | brownish, usually spiny, sometimes prominently cristate, rarely finely reticulate. |
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Basal | pinnae 8–18 × 2–3 cm, equilateral, incised more than 3/4 distance to costae. |
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x | = 41. |
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Ctenitis |
Ctenitis submarginalis |
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Habitat | Cypress swamps, hammocks, old forested spoil banks | |||||
Elevation | 0 m (0 ft) | |||||
Distribution |
Nearly worldwide in the tropics |
FL; LA; e Mexico; s Mexico; Central America; West Indies in Hispaniola; South America to Uruguay |
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Discussion | Species ca. 100 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Combining authorship of the accepted name sometimes has been incorrectly attributed to E. B. Copeland (1947). The Louisiana population of Ctenitis submarginalis occurs more than 960 km from populations in Florida and represents the northernmost locality for the species (G. P. Landry and W. D. Reese 1991). Unlike most ferns in North America, the plants in the Louisiana population are nonseasonal, producing leaves and sori throughout the year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Dryopteris subg. C. | Polypodium submarginale, Dryopteris submarginalis | ||||
Name authority | (C. Christensen) C. Christensen: in Verdoorn et al., Man. Pteridol. 543. (1938) | (Langsdorff & Fischer) Ching: Sunyatsenia 5: 250. (1940) | ||||
Web links |