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comb fern, lacefern

Habit Plants generally terrestrial.
Stems

erect to obliquely ascending, stolons absent.

Leaves

monomorphic, evergreen or dying back in winter.

Petiole

2/3 to equaling length of blade, base not swollen;

vascular bundles more than 3, arranged in an arc, ± round in cross section.

Blade

lanceolate to deltate, 1–4-pinnate-pinnatifid, gradually reduced distally to confluent, pinnatifid apex, herbaceous.

Pinnae

not articulate to rachis, segment margins nearly entire to crenulate, ciliate;

proximal pinnae not reduced, sometimes basal pair much the longest, sessile to petiolulate, equilateral or inequilateral with basiscopic side more developed (pinnules noticeably longer);

costae adaxially rounded or flat, not grooved;

indument of linear to lanceolate scales and often multicellular glandular hairs abaxially, of multicellular reddish hairs adaxially.

Veins

free, simple or forked.

Sori

in 1 row between midrib and margin, round;

indusia round-reniform, attached at narrow sinus, sometimes small or seemingly absent, persistent or caducous.

Spores

brownish, usually spiny, sometimes prominently cristate, rarely finely reticulate.

x

= 41.

Ctenitis

Distribution
from USDA
Nearly worldwide in the tropics
[BONAP county map]
Discussion

Species ca. 100 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Blades ovate-lanceolate, 2-4-pinnate-pinnatifid; basal pinnae inequilateral, elongate basiscopically.
C. sloanei
1. Blades oblong or narrowly lanceolate, 1-pinnate-pinnatifid; basal pinnae equilateral.
C. submarginalis
Source FNA vol. 2. Author: Robbin C. Moran.
Parent taxa Dryopteridaceae
Subordinate taxa
C. sloanei, C. submarginalis
Synonyms Dryopteris subg. C.
Name authority (C. Christensen) C. Christensen: in Verdoorn et al., Man. Pteridol. 543. (1938)
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